| 词条 | Notholaena standleyi |
| 释义 |
| image = Notholaena standleyi.jpg | image_caption = Notholaena standleyi. Standley's cloak fern from Phoenix Mountains Preserve, Maricopa County, Arizona, USA. | genus = Notholaena | species = standleyi | authority = Maxon }} Notholaena standleyi, also known as star cloak fern and Standley's cloak fern, is a fern that is native to the United States and Mexico. It is a member of the genus Notholaena, which is part of the subfamily Cheilanthoideae of family Pteridaceae. It is distinguished by the pentagonal shape formed by the blades of its frond, a property other members of Notholaena do not possess. TaxonomyNotholaena standleyi was named after Paul C. Standley, Assistant Curator of the United States National Herbarium, who studied plants in southwestern United States. First described as Notholaena candida var. quinquefidopalmata in 1864, it was elevated to a species as Notholaena hookeri in 1879 by D. C. Eaton. However, this name had already been used to describe another fern, and it was renamed by William R. Maxon in 1915. He named it after Standley, who had engaged in a great deal of field study in the southwestern United States. Standley was also collaborating with E. O. Wooton to develop a comprehensive Flora of New Mexico at the time.[1]DistributionNotholaena standleyi is indigenous to southwestern United States and parts of north and central Mexico. It specifically grows in Arizona, Colorado, New Mexico, Oklahoma, and Texas in the USA.[2]Habitat and EcologyNotholaena standleyi is a perennial species that typically grows in desert regions at elevations from 300 to 2100 m.[3] It is found on rocky hillsides, usually in the crevices created by limestone and granite boulders that provide the partial shade the plant prefers. During periods of drought, the frond may curl and become brown until water is available, an adaptation to the semi-arid environments it inhabits.[4] At lower elevations, it sometimes grows alongside Notholaena californica.[5]It takes 13 days to germinate and sporulates from late spring to fall.[3] It has the same life cycle as a typical fern, the sporophyte produces haploid spores by meiosis, which grows into a haploid gametophyte. The gametophyte produce gametes by mitosis. The fertilized egg when develop into the sporophyte by mitosis. The spore is transported by air and water. MorphologyIndividuals of this species have brown petioles and green fronds. The ferns are erect and grow to a height of {{convert|25|cm|in}}. The petioles grow to a length of {{convert|5|to|13|cm|in}}. The individual fronds are usually {{convert|3|to|7|cm|in}} wide as well as long.[5] The frond is light green when young, becoming darker green as it matures, and usually contains five blades arranged in a pentagonal fashion. The blades are bipinnately or tripinnately compound, and each of them is triangular and pointed. The abaxial side of the frond is covered by powdery farina, with a function that is still unclear.[6] The color of the farina differs between species growing in different geographic locations. These are referred to as chemotypes, of which there are four. These chemotypes also have different number of complete chromosome sets. Gold and yellow farina are diploids, pallid and yellow/green are tetraploids.[7] Its prothallus contains glandular trichomes. It usually has 32 spores per sporangium, but many with only 16 have been observed, produced from eight 2n mother cells.[8] It is diploid and has 60 chromosomes in its root tip cells.[9] UsageThe Seri Indians made tea from the fronds to promote fertility in women. The species was also believed to have supernatural value; the Seri carried in a small cloth bag to bring good luck to the bearer.[4] It has been studied in potential herpes treatments, because of its antiviral activity. Notholaenic acid, isolated from the species, has shown anti-HSV-1 activity at high concentrations (25 μg/disk).[10] {{-}}References1. ^{{cite journal|last=Maxon|first=William R.|authorlink=William Ralph Maxon|title=Note on American Ferns: IX.|journal=American Fern Journal|date=Jan–Mar 1915|volume=5|issue=1|pages=1|url=https://books.google.com/books?id=rRvyAAAAMAAJ&pg=PA1|accessdate=20 April 2012|doi=10.2307/1544458|jstor=1544458}} 2. ^{{cite web|title=Notholaena standleyi Maxon|url=http://plants.usda.gov/java/profile?symbol=NOST|work=PLANTS Profile|publisher=USDA|accessdate=17 April 2012}} 3. ^1 {{cite book|last=Flora of North America Editorial Committee|title=Flora of North America: Pteridophytes and Gymnosperms|year=1993|publisher=Oxford University Press|isbn=978-0195082425|pages=148–149|url=https://books.google.com/books?id=VnlBQMJOi-UC&pg=PA148}} 4. ^1 {{cite book|last=Austin|first=Daniel|title=Baboquivari Mountain Plants: Identification, Ecology, and Ethnobotany|year=2010|publisher=University of Arizona Press|isbn=978-0816528370|page=232|url=https://books.google.com/?id=nnFVFdp0dWIC&pg=PA232&lpg=PA232&dq=cheilanthes+standleyi+usage#v=onepage&q=cheilanthes%20standleyi%20usage&f=false}} 5. ^1 {{cite book | last=Felger | first=Richard Stephen | title=Flora of the Gran Desierto and Río Colorado of Northwestern Mexico | publisher=The University of Arizona Press | year=2000 | isbn=978-0-8165-2044-2 | page=47 | url=https://books.google.com/books?id=ICgHfrVNLUUC&pg=PA47}} 6. ^{{cite book|last1=Yarborough |first1=Sharon C. | last2=Powell | first2=A. Michael|title=Ferns and Fern Allies of the Trans-Pecos and Adjacent Areas|year=2002|publisher=Texas Tech University Press|isbn=978-0896724761|page=67|url=https://books.google.com/books?id=Gx0SucseK1QC&pg=PA67}} 7. ^{{cite web|last=Rothfels|first=Carl|title=Notholaena standleyi Maxon 1915. Northern Desert Star Cloakfern|date=23 December 2008|url=http://tolweb.org/Notholaena_standleyi/133579/2008.12.23|publisher=The Tree of Life Web Project|accessdate=2012-05-01}} 8. ^{{cite journal|last=Seigler|first=David S.|author2=E. Wollenweber |title=Chemical Variation in Notholaena standleyi|journal=American Journal of Botany|date=May–June 1983|volume=70|issue=5|pages=790–798|jstor=2443133|doi=10.2307/2443133}} 9. ^{{cite journal|last=Knobloch|first=Irving W.|author2=William Tai |author3=Thamby A. Ninan |title=The Cytology of Some Species of the Genus Notholaena|journal=American Journal of Botany|date=Jan 1973|volume=60|issue=1|pages=92–95|jstor=2441329|doi=10.2307/2441329}} 10. ^{{cite journal|first1=Kenneth L. |last1=Rinehart |first2=Tom G. |last2=Holt |first3=Nancy L. |last3=Fregeau |first4=Paul A. |last4=Keifer |first5=George Robert |last5=Wilson |first6=Thomas J., Jr. |last6=Perun |first7=Ryuichi |last7=Sakai |first8=Anthony G. |last8=Thompson |first9=Justin G. |last9=Stroh |first10=Lois S. |last10=Shield |first11=David S. |last11=Seigler |first12=Li H. |last12=Li |first13=David G. |last13=Martin |first14=Cornelis J. P. |last14=Grimmelikhuijzen |first15=Gerd |last15=Gäde|title=Bioactive Compounds from Aquatic and Terrestrial Sources|journal=Journal of Natural Products|date=July 1990|volume=53|issue=4 |pages=771–792|doi=10.1021/np50070a001}} External links{{Commons category|Notholaena standleyi}}{{Taxonbar|from=Q7062960}} 10 : Notholaena|Ferns of Mexico|Ferns of the United States|Flora of the Southwestern United States|Flora of Northwestern Mexico|Flora of Northeastern Mexico|Flora of the Chihuahuan Desert|Flora of the Sonoran Deserts|Plants described in 1915|Pteridophyta of the Americas |
| 随便看 |
|
开放百科全书收录14589846条英语、德语、日语等多语种百科知识，基本涵盖了大多数领域的百科知识，是一部内容自由、开放的电子版国际百科全书。