| 词条 | Ulmaceae |
| 释义 |
|image = Ulmus laciniata.jpg |image_caption = Ulmus laciniata Morton Arboretum acc. 180-84-1 |taxon = Ulmaceae |authority = Mirb. 1815 |type_species = Ulmus |type_species_authority = L. 1753 |subdivision_ranks = Genera |subdivision =
| range_map = Ulmaceae Distribution.svg | range_map_caption = The range of Ulmaceae. | synonyms = * Samaracaceae Dulac | synonyms_ref = [1] }} The Ulmaceae ({{IPAc-en|ʌ|l|ˈ|m|eɪ|s|i}}) are a family of flowering plant that includes the elms (genus Ulmus), and the zelkovas (genus Zelkova).[1] Members of the family are widely distributed throughout the north temperate zone, and have a scattered distribution elsewhere except for Australasia.[2][3] The family was formerly sometimes treated to include the hackberries, (Celtis and allies), but analysis by the Angiosperm Phylogeny Group members suggests that these genera are better placed in the related family Cannabaceae. The circumscription included in the taxobox is the one suggested by P. Stevens on his Missouri Botanical Garden Angiosperm Phylogeny Website and includes information from the Royal Botanic Gardens, Kew Vascular Plant Families and Genera list.[3] It generally is considered to include ca 7 genera and about 45 species.[4] Some classifications also include the genus Ampelocera.[5] DescriptionThe family is a group of evergreen or deciduous trees and shrubs with mucilaginous substances in leaf and bark tissue. Leaves are usually alternate on the stems. The leaf blades are simple (not compound), with entire (smooth) or variously toothed margins, and often have an asymmetrical base. The flowers are small and either bisexual or unisexual.[6] The fruit is an indehiscent samara, nut, or drupe. Ulmus provides important timber trees mostly for furniture, and U. rubra, the slippery elm, is a medicinal plant known for the demulcent property of its inner bark. Planera aquatica is also a timber species. Planera, Ulmus, and Zelkova are all grown as ornamental trees. PhylogenyModern molecular phylogenetics suggest the following relationships:[5][6][7][8][9] {{Clade| style=line-height:75%;|1={{clade |1=Cannabaceae (outgroup) |label2=Ulmaceae |2={{clade |1={{clade |1=Holoptelea |2=Ampelocera |2={{clade |1=Hemiptelea |2={{clade |1=Zelkova |2=Ulmus }} }} }} }} }} References1. ^{{cite journal | last = Denk | first = T | author2 = GW Grimm | title = Phylogeny and biogeography of Zelkova (Ulmaceae sensu stricto) as inferred from leaf morphology, ITS sequence data and the fossil record | journal = Bot J Linn Soc | volume = 147 | issue = 2 | pages = 129–157 | date = February 2005 | doi = 10.1111/j.1095-8339.2005.00354.x}} {{Taxonbar|from=Q156311}}2. ^1 {{cite web|last = Watson|first = L.|author2=Dallwitz, M. J. |title = The Families of Flowering Plants: Ulmaceae Mirb.|date = 1992 onwards|url = http://delta-intkey.com/angio/www/ulmaceae.htm|accessdate = 21 November 2006 }} 3. ^1 {{cite web|last = Stevens|first = P|title = Angiosperm Phylogeny Website|publisher = Missouri Botanical Garden|date = 2001 onwards|url = http://www.mobot.org/mobot/research/APweb/welcome.html|accessdate = 21 November 2006 }} 4. ^{{cite journal | author = Christenhusz, M. J. M., and Byng, J. W. | year = 2016 | title = The number of known plants species in the world and its annual increase | journal = Phytotaxa | volume = 261 | pages = 201–217 | url = http://biotaxa.org/Phytotaxa/article/download/phytotaxa.261.3.1/20598 | doi = 10.11646/phytotaxa.261.3.1 | issue = 3 }} 5. ^1 {{cite journal|last = Ueda|first = Kunihiko |author2=K Kosuge |author3=H Tobe|title = A molecular phylogeny of Celtidaceae and Ulmaceae (Urticales) based on rbcL nucleotide sequences |journal = Journal of Plant Research |volume = 110|issue = 2|pages = 171–178|date=June 1997|doi = 10.1007/BF02509305}} 6. ^1 {{Citation |last=Sytsma |first=Kenneth J. |last2=Morawetz |first2=Jeffery |last3=Pires |first3=J. Chris |last4=Nepokroeff |first4=Molly |last5=Conti |first5=Elena |last6=Zjhra |first6=Michelle |last7=Hall |first7=Jocelyn C. |last8=Chase |first8=Mark W. |year=2002 |title=Urticalean rosids: Circumscription, rosid ancestry, and phylogenetics based on rbcL, trnL–F, and ndhF sequences |journal=Am J Bot |volume=89 |issue=9 |pages=1531–1546 |doi=10.3732/ajb.89.9.1531 |lastauthoramp=yes |pmid=21665755}} 7. ^{{cite journal | authors = Zavada MS, Kim M. | year = 1996 | title = Phylogenetic analysis of Ulmaceae | journal = Plant Syst Evol | volume = 200 | issue = 1 | pages = 13–20 | doi = 10.1007/BF00984745}} 8. ^{{cite conference | url = http://www.2012.botanyconference.org/engine/search/index.php?func=detail&aid=316 | title = Fossils, biogeography and dates in an expanded phylogeny of Ulmaceae | authors = Neubig K, Herrera F, Manchester S, Abbott JR. | date = July 7–11, 2012 | publisher = Botanical Society of America | location = St. Louis, Missouri | conference = Botany 2012: Annual Meeting of the Botanical Society of America in Columbus, Ohio, USA | id = Abstract 316}} 9. ^{{cite journal | authors = Sun M, Naeem R, Su J-X, Cao Z-Y, J. Burleigh G, Soltis PS, Soltis DE, Chen Z-D. | year = 2016 | title = Phylogeny of the Rosidae: A dense taxon sampling analysis | journal = Journal of Systematics and Evolution | volume = 54 | issue = 4 | pages = 363–391 | doi = 10.1111/jse.12211}} 2 : Ulmaceae|Rosid families |
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