| 词条 | Verbena | |||||
| 释义 |
|image = Verbena bonariensis1.jpg |image_caption = Purpletop vervain, Verbena bonariensis |taxon = Verbena |authority = L.[1] |type_species = Verbena officinalis |type_species_authority = L. |subdivision_ranks = Species |subdivision =
|synonyms = Shuttleworthia Meisn.[1] }} Verbena ({{IPAc-en|v|ər|ˈ|b|iː|n|ə}},[3] vervain) is a genus in the family Verbenaceae. It contains about 250 species of annual and perennial herbaceous or semi-woody flowering plants. The majority of the species are native to the Americas and Asia. DescriptionThe leaves are usually opposite, simple, and in many species hairy, often densely so. The flowers are small, with five petals, and borne in dense spikes. Typically some shade of blue, they may also be white, pink, or purple, especially in cultivars. The genus can be divided into a diploid North American and a polyploid South American lineage, both with a base chromosome number of seven. The European species is derived from the North American lineage. It seems that verbena as well as the related mock vervains (Glandularia) evolved from the assemblage provisionally treated under the genus name Junellia; both other genera were usually included in the Verbenaceae until the 1990s.[4] Intergeneric chloroplast gene transfer by an undetermined mechanism – though probably not hybridization – has occurred at least twice from vervains to Glandularia, between the ancestors of the present-day South American lineages and once more recently, between V. orcuttiana or V. hastata and G. bipinnatifida. In addition, several species of verbena are of natural hybrid origin; the well-known garden vervain has an entirely muddy history. The relationships of this close-knit group are therefore hard to resolve with standard methods of computational phylogenetics.[5] CultivationSome species, hybrids and cultivars of verbena are used as ornamental plants. They are drought-resistant, tolerating full to partial sun, and enjoy well-drained, average soils. Plants are usually grown from seed. Some species and hybrids are not hardy and are treated as half-hardy annuals in bedding schemes.[6] They are valued in butterfly gardening in suitable climates, attracting Lepidoptera such as the Hummingbird hawk-moth, Chocolate albatross, or the Pipevine swallowtail, and also hummingbirds, especially V. officinalis, which is also grown as a honey plant. The hybrid cultivars "Silver Anne"[7] and "Sissinghurst"[8] have gained the Royal Horticultural Society's Award of Garden Merit. Pests and diseasesFor some verbena pathogens, see List of verbena diseases. Cultivated verbenas are sometimes parasitized by Sweet potato whitefly (Bemisia tabaci) and spread this pest to other crops. UsesAlthough verbena ("vervain") has been used in herbalism and traditional medicine, usually as an herbal tonic, there is no high-quality evidence for its effectiveness.[9] Verbena has been listed as one of the 38 plants used to prepare Bach flower remedies,[10] a kind of alternative medicine promoted for its effect on health. According to Cancer Research UK, "essence therapists believe that using essences can help to increase your mental, emotional and spiritual wellbeing. However, essences are not used to prevent, control, or cure cancer or any other physical condition."[11] The essential oil of various species, mainly common vervain, is traded as "Spanish verbena oil". Considered inferior{{citation needed|date=March 2018}} to oil of lemon verbena (Aloysia triphylla) in perfumery,[9] it is of some commercial importance for herbalism. In culture{{see also|Verbena (disambiguation)}}Verbena has long been associated with divine and other supernatural forces. It was called "tears of Isis" in ancient Egypt, and later called "Hera's tears". In ancient Greece it was dedicated to Eos Erigineia. In the early Christian era, folk legend stated that V. officinalis was used to staunch Jesus' wounds after his removal from the cross. It was consequently called "holy herb" or (e.g. in Wales) "Devil's bane".{{citation needed|date=December 2013}} Vervain flowers are engraved on cimaruta, Italian anti-stregheria charms.{{citation needed|date=December 2013}} In the 1870 The History and Practice of Magic by "Paul Christian" (Jean-Baptiste Pitois) it is employed in the preparation of a mandragora charm.[12] The book also describes its antiseptic capabilities (p. 336), and use as a protection against spells (pp. 339, 414).[13] While common vervain is not native to North America, it has been introduced there and for example the Pawnee have adopted it as an entheogen enhancer and in oneiromancy (dream divination), much as Calea zacatechichi is used in Mexico. The generic name is the Latin term for a plant sacred to the ancient Romans.[14][15] Pliny the Elder describes verbena presented on Jupiter altars; it is not entirely clear if this referred to a verbena rather than the general term for prime sacrificial herbs.{{Verify source|date=November 2007}} The common names of verbena in many Central and Eastern European languages often associate it with iron. These include for example the Dutch {{lang|nl|IJzerhard}} ("iron-hard"), Danish {{lang|da|Læge-Jernurt}} ("medical ironwort"), German {{lang|de|Echtes Eisenkraut}} ("true ironherb"), Slovak {{lang|sk|Železník lekársky}} ("medical ironherb"), and Hungarian {{lang|mg|vasfű}} ("iron grass"). An indeterminate vervain{{Verify source|date=November 2007}} is among the plants on the eighth panel of the New World Tapestry (Expedition to Cape Cod).{{citation needed|date=December 2013}} In the Victorian language of flowers, verbena held the dual meaning of enchantment and sensibility.[16] SpeciesVerbena has the following subtaxa.[17] See also Aloysia, Glandularia and Junellia for species formerly placed here. {{div col|colwidth=24em}}
GalleryReferences1. ^1 {{cite web |publisher=United States Department of Agriculture |work=Germplasm Resources Information Network |url=http://www.ars-grin.gov/cgi-bin/npgs/html/genus.pl?12680 |title=Genus: Verbena L. |date=2004-01-29 |accessdate=2011-08-29}} 2. ^1 {{cite web |publisher=United States Department of Agriculture |work=Germplasm Resources Information Network |url=http://www.ars-grin.gov/cgi-bin/npgs/html/splist.pl?12680 |title=GRIN Species Records of Verbena |accessdate=2011-08-29}} 3. ^Sunset Western Garden Book, 1995:606–607 4. ^{{cite journal |author=S. M. Botta, S. Martinez & M. E. Mulguta de Romero |year=1995 |title=Novedades nomenclaturales en Verbenaceae |trans-title=Nomenclatural revisions in Verbenaceae |journal=Hickenia |volume=2 |pages=127–128}} 5. ^{{cite journal |author1=Yao-Wu Yuan |author2=Richard G. Olmstead |lastauthoramp=yes |year=2008 |title=A species-level phylogenetic study of the Verbena complex (Verbenaceae) indicates two independent intergeneric chloroplast transfers |journal=Molecular Phylogenetics and Evolution |volume=48 |issue=1 |pages=23–33 |doi=10.1016/j.ympev.2008.04.004 |pmid=18495498}} 6. ^{{cite book|title=RHS A-Z encyclopedia of garden plants|year=2008|publisher=Dorling Kindersley|location=United Kingdom|isbn=978-1405332965|page=1136}} 7. ^{{cite web|url=http://apps.rhs.org.uk/plantselector/plant?plantid=5445 |title=RHS Plant Selector Verbena 'Silver Anne' (G) AGM / RHS Gardening |publisher=Apps.rhs.org.uk |accessdate=2013-05-06}} 8. ^{{cite web|url=http://apps.rhs.org.uk/plantselector/plant?plantid=4286 |title=RHS Plant Selector Verbena 'Sissinghurst' (G) AGM / RHS Gardening |publisher=Apps.rhs.org.uk |accessdate=2013-05-06}} 9. ^1 {{cite web|title=Vervain|url=https://www.drugs.com/npp/vervain.html|publisher=Drugs.com|accessdate=7 March 2018|date=2009}} 10. ^{{cite book|author=D. S. Vohra|title=Bach Flower Remedies: A Comprehensive Study|url=https://books.google.com/books?id=icG8onA0ys8C&pg=PR3|accessdate=2 September 2013|date=1 June 2004|publisher=B. Jain Publishers|isbn=978-81-7021-271-3|page=3}} 11. ^{{cite web|url=http://www.cancerresearchuk.org/about-cancer/cancer-in-general/treatment/complementary-alternative-therapies/individual-therapies/essence-therapy|title=Flower remedies|publisher=Cancer Research UK|accessdate=11 November 2016}} 12. ^Pitois, Christian (1952). [https://books.google.com/books?id=DXIKAQAAIAAJ&focus=searchwithinvolume&q=mandragora+verbena The History and Practice of Magic, Volume 2], Forge Press. p. 402. 13. ^Pitois, (1952) [https://books.google.com/books?id=DXIKAQAAIAAJ&focus=searchwithinvolume&q=verbena p. 336, 339, 414] 14. ^{{Cite book |url=https://books.google.com/books?id=2ndDtX-RjYkC |title=CRC World Dictionary of Plant Names |volume=4 R-Z |year=2000 |first=Umberto |last=Quattrocchi |publisher=Taylor & Francis US |isbn=978-0-8493-2678-3 |page=2787}} 15. ^{{cite book |url=https://books.google.com/books?id=NJ6PyhVuecwC |title=The Names of Plants |first=D. |last=Gledhill |edition=4 |publisher=Cambridge University Press |isbn=978-0-521-86645-3 |year=2008 |page=399}} 16. ^{{Cite web|url=http://www.languageofflowers.com/flowermeaning.htm|title=Language of Flowers - Flower Meanings, Flower Sentiments|website=www.languageofflowers.com|access-date=2016-11-26}} 17. ^{{cite web|author1=|title=Verbena|url=https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Tree&id=22002&lvl=3&keep=1&srchmode=1&unlock|website=NCBI taxonomy|publisher=National Center for Biotechnology Information|accessdate=21 June 2018|location=Bethesda, MD|language=English|format=HTML }} External links{{Commons category}}{{Wikispecies}}{{Medicinal herbs & fungi}}{{Taxonbar|from=Q165290}} 4 : Verbena|Garden plants|Lamiales genera|Medicinal plants |
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