| 词条 | Amanita gemmata |
| 释义 |
| image = Amanita gemmata von hms.jpg | image_width = 234px | image_caption = In different stages of development | regnum = Fungi | divisio = Basidiomycota | classis = Agaricomycetes | ordo = Agaricales | familia = Amanitaceae | genus = Amanita | species = A. gemmata | binomial = Amanita gemmata | binomial_authority = (Fr.) Bertill. (1866) | synonyms = {{collapsible list|title=Species synonymy[1][2][3] |Agaricus gemmatus Fr. (1838) |Agaricus adnatus W.G.Sm. (1870) |Amanita gemmata (Fr.) Gillet (1874) |Amanita junquillea Quél. (1876) |Amanita gemmata f. amici Gillet (1884) |Amanita muscaria var. gemmata (Fr.) Quél. (1886) |Amanitopsis adnata (W.G.Sm.) Sacc. (1887) |Amanitopsis gemmata (Fr.) Sacc. (1887) |Amanita adnata (W.G.Sm.) Sacc. (1925) |Amanita junquillea var. exannulata J.E.Lange (1935) |Amanitaria gemmata (Fr.) E.-J.Gilbert (1940) |Venenarius gemmatus (Fr.) Murrill (1948) }}{{mycomorphbox | name = Amanita gemmata | whichGills = adnate | whichGills2 = adnexed | capShape = convex | capShape2 = flat | hymeniumType=gills | stipeCharacter=ring | ecologicalType=mycorrhizal | sporePrintColor=white | howEdible=poisonous | howEdible2=deadly }}{{stack end}} Amanita gemmata, commonly known as the gemmed Amanita or the jonquil Amanita, is an agaric mushroom of the family Amanitaceae and genus Amanita. The fruit body has a cap that is a dull to golden shade of yellow, and typically {{convert|2.5|–|12|cm|in|1|abbr=on}} in diameter. The cap surface is sticky when moist, and characterized by white warts, which are easily detached. It is initially convex, and flattens out when mature. The flesh is white and does not change colour when cut. Gills are white and closely spaced. The stem is pale yellow, and measures {{convert|4|–|12|cm|in|abbr=on}} long by {{convert|0.5|–|1.9|cm|in|1|abbr=on}} thick. The partial veil that covers the young fruit body turns into the ring on the stem at maturity. The spore print is white, while the spores are roughly elliptical, and measure 8–10 by 6.5–7.5 µm. This species is a mycorrhizal fungus, widespread in Europe. It can grow either singly, scattered, or in groups. It prefers habitats like coniferous and mixed forests and alongside paths, where it fruits in summer and fall. It is a toxic mushroom that has muscarine, the same toxin as Amanita muscaria and Amanita pantherina as well as many species in the Clitocybe and Inocybe genera. It is often confused with various other European species. A. gemmata resembles the false death cap, tawny grisette and panther cap mushrooms. Its cap is brighter in color than the former, and more yellow than the latter two. Taxonomy and phylogeny{{cladogram|align=left|title=|clade={{clade |style=font-size:75%;line-height:75% |label1= |1={{clade |1={{clade |1={{clade |1=Amanita muscaria |2={{clade |1=Amanita roseitincta |2=Amanita gemmata }} }} |2=Amanita farinosa |2={{clade |1={{clade |1={{clade |1=Amanita ceciliae |2=Amanita fulva |2=Amanita vaginata |2=Amanita jacksonii }} }} }} |caption=Phylogeny and relationships of A. gemmata and related species based on combined analyses of nLSU-DNA and mtSSU-DNA sequences.[4] }} The species was first described scientifically by Swedish mycologist and botanist Elias Magnus Fries as Agaricus gemmatus in 1838.[5] It was transferred to the genus Amanita in 1866 by the French statistician Louis Bertillon.[6] The species has been transferred to several genera in its history, resulting in a number of synonyms, including Amanita muscaria var. gemmata (1886, Lucien Quélet), Amanitopsis gemmata (1887, Pier Andrea Saccardo), Amanitaria gemmata (1940, Jean-Edouard Gilbert), and Venenarius gemmatus (1948, William Murrill).[1] Amanita authority Rodham E. Tulloss considers A. amici (published by Claude Casimir Gillet in 1891) to be synonymous with A. gemmata, as the macroscopic characteristics of the former fall within the limits of the range expected for the latter.[3] Within the genus Amanita, A. gemmata is classified in subgenus Amanita, section Amanita, subsection Gemmatae, and series Gemmatae.[3] Tulloss places the species in a stirps (an informal ranking above species level) with A. russuloides and A. viscidolutea.[10] Some mycologists believe that A. gemmata is not different from A. russuloides.[11] Two molecular studies based on sequences of the large ribosomal subunit RNA gene (nLSU-rDNA) and the mitochondrial small ribosomal subunit RNA gene (mtSSU-rDNA) show that A. gemmata is part of a clade within Amanita with its close relatives A. muscaria, A. farinosa and A. roseitincta.[4][13] The mushroom is commonly known as the "gemmed Amanita", the "jonquil Amanita",[14] or the "European gemmed Amanita".[3] DescriptionThe fruit bodies are colored yellow overall. The fresh cap, ranging in color from dull creamy yellow to golden yellow to buff, is sticky when moist. White warts adorn the cap surface, but they are usually flimsy and easily washed away by rain. They are placed randomly, but tend to be more concentrated in the center. The cap is typically {{convert|2.5|–|12|cm|in|1|abbr=on}} in diameter, and initially convex before flattening out in maturity. The flesh is white, and shows no change when sliced. The gills are adnate to adnexed, and white; they are close together, with little intervening space. The pale yellowish stem is {{convert|4|–|12|cm|in|abbr=on}} long by {{convert|0.5|–|1.9|cm|in|1|abbr=on}} thick, and either roughly equal in width throughout, or slightly thicker at the base. Young mushrooms have a membranous partial veil extending from the upper stem to the cap margin; as the mushroom grows, the partial veil tears to leave a flimsy, skirt-like, easily lost ring on the stem. At the base of the stem is a white volva (a remnant of the universal veil that covered the immature mushroom) that usually forms a small, free rim. Spore prints are white.[14] There is no distinctive odour.[18] Microscopic characteristicsAmanita gemmata has ellipsoid to broadly ellipsoid spores measuring 8–10 by 6.5–7.5 µm with an average Q-ratio (the fraction of length/width) of 1.35; they are not amyloid. The spores are smooth, thin-walled, and they contain one to several small oil droplets.[3] The basidia (spore-bearing cells of the hymenium) are usually four-spored, club shaped, and measure 30–40 by 8–11 µm.The gill tissue is divergent, meaning that the cells are more or less parallel near the center of the gill, but bend outwards near the end of the gill. The hyphae in this tissue are cylindrical to inflated, thin walled, hyaline (translucent) to yellowish, and measure 2.2–9 µm wide; the hyphae in the central strand are narrower and typically cylindrical. The hyphae of the subhymenium (a layer of tissue directly under the hymenium) are interwoven. These hyphae are branched, cylindrical to slightly inflated, hyaline, and 6–9 µm wide. The hyphae of the cap cuticle are filamentous, interwoven, and radially arranged. They are cylindrical, 2.7–4 µm wide, thin-walled, hyaline to yellowish, and gelatinize when mounted in potassium hydroxide. The cap tissue is also interwoven, with hyphae that are cylindrical to somewhat inflated, 3.7–14.6 µm wide, thin-walled, branched, and hyaline to yellowish. Caulocystidia are abundant on the apex of the stem; they are club-shaped to cylindrical, thin-walled, hyaline, and measure 3–9 µm wide. The annulus tissue comprises interwoven cylindrical hyphae measuring 3–9 µm wide. Sphaerocysts (inflated, spherical cells) are also present in the annulus tissue; they are club shaped to ellipsoidal, with dimensions of 29–55 by 30–70 µm. The warts on the cap surface (remnants of the universal veil) comprise loosely interwoven cylindrical to inflated thin-walled hyphae that are 3.5–8 µm wide. Sphaerocysts in this tissue are 58.5–70.2 by 17.5–40 µm, ellipsoidal, and hyaline. The volval tissue is interwoven, with cylindrical, hyaline hyphae that are 4.4–7.3 µm wide. The sphaerocysts here are ellipsoidal to roughly spherical, hyaline, and measure 35–70 by 20–35 µm. In A. gemmata, where they are most abundant in the region just below the cap cuticle, these refractive cells are scattered, and have a width of 3.7–6 µm. Clamp connections are rare in the hyphae of A. gemmata; they are present in the annulus, gill tissue, subhymenium, and cap tissue. Similar speciesThere are numerous species in North America that resemble A. pantherina.[22] In 2005, mycologist Rod Tulloss described Amanita aprica, a species that has been confused with A. gemmata several times in the past.[23] According to mycologists Pierre Neville and Serge Poumarat, the Mediterranean species A. amici (synonymous with A. gemmata f. amici[24]) is similar in appearance to A. gemmata but is larger. According to Tulloss however, their measurements of the cap and stem dimensions of A. amici fell within the range expected for A. gemmata, and for this reason, the two taxa should be considered conspecific.[3] Neville and Poumarat suggest that the name A. gemmata still persists for Mediterranean collections because of its frequent historical misapplication to the native Italian species A. gioiosa, which had not been described as a distinct species until 2004.[3] A. orientigemmata, a mushroom ranging from Japan to China, is a lookalike, but has clamps, unlike A. gemmata. Other differences between the two species include the slightly smaller spores of A. orientigemmata, and differences in the microstructure of the cap warts.[27] ToxicityToxicity is suspected to be due to the presence of muscimol and ibotenic acid.[28] Generally, symptoms of poisoning appear within three hours of ingestion of the mushroom as visual hallucinations, nausea, vomiting, stomach pain, diarrhea, irregular and slow heart beat and agitation. Severe cases involving coma, convulsions, or death are extremely rare.[29] Ecology, habitat and distributionAmanita gemmata is a mycorrhizal fungus, meaning it forms a mutually beneficial relationship with the roots of compatible host plants. Through the association, the plant provides the fungus with a carbon source, and the fungus provides the plant with several benefits such as nutrients and protection from pathogens. Largent and collaborators (1980) document mycorrhizal associations of A. gemmata with Manzanita (Arctostaphylos spp.) and Lodgepole Pine (Pinus contorta),[30] and Nieto and Carbone with Maritime Pine (Pinus pinaster) in Spain.[31] The fungus favours sandy and slightly acidic soils, and is often found in association with Norway Spruce (Picea abies). The mushroom grows either singly, scattered, or in groups in coniferous and mixed forests, especially along paths and roads.[14][33]The species is distributed in areas of Asia and Europe. It is widely distributed (as a species cluster, not the actual Amanita gemmata) in North America,[14] where it has been found as far south as Ixtlán de Juárez, Mexico.[35] The species has been reported from the Dominican Republic.[36] In South America, it is known from Chile.[37] In Asia, the mushroom has been collected from Iran[38] and China.[39] See also{{Portal|Fungi}}
References1. ^1 2 3 4 {{cite book |author =Arora, David |authorlink=David Arora |title=Mushrooms Demystified: A Comprehensive Guide to the Fleshy Fungi |publisher=Ten Speed Press |location=Berkeley, California |year=1986 |page=281 |isbn=978-0-89815-169-5}} [1][2][3][4][5][6][7][8][9][10][11][12][13][14][15][16][17][18][19][20][21][22][23][24][25]2. ^1 {{cite book |author =Dechambre, Amédée |title=Dictionnaire Encyclopédique des Sciences Médicales |year=1866 |volume=1 |issue=3 |pages=491–511 |publisher=Masson |location=Paris, France |language=French}} 3. ^1 {{cite book |author1=Bresinsky, Andreas |author2=Besl, Helmut |title=A Colour Atlas of Poisonous Fungi: A Handbook for Pharmacists, Doctors, and Biologists |publisher=Manson Publishing |location=London, UK |year=1989 |pages=110–111 |isbn=978-0-7234-1576-3 |url=https://books.google.com/books?id=EIcQGsZ2kksC&lpg=PA110&dq=amanita%20gemmata&pg=PA110}} 4. ^1 {{cite book |author =Fries, Elias Magnus |authorlink=Elias Fries |title=Epicrisis systematis mycologici: seu synopsis hymenomycetum |year=1838 |publisher=Typographia Academica |location=Uppsala, Sweden |page=12 |url=https://books.google.com/books?id=bYdIAAAAYAAJ&pg=PA12 |language=Latin}} 5. ^1 {{cite journal |author =Gallart, C. Augusto Rodriguez |title=Study of micromycetes from the Dominican Republic: Part III |journal=Moscosoa |year=1997 |volume=9 |pages=145–153 |language=Spanish |issn=0254-6442}} 6. ^1 {{cite book |author1=Gilbert, John |author2=Şenyuva, Hamide |title=Bioactive Compounds in Foods |year=2008 |publisher=Blackwell Publishing |location=Oxford, UK |isbn=978-1-4051-5875-6 |page=119 |url=https://books.google.com/books?id=-2myBYRAGogC&pg=PA119}} 7. ^1 {{cite web |author =Kuo, Michael |date=March 2006 |title=Amanita gemmata species group |url=http://www.mushroomexpert.com/amanita_gemmata.html |publisher=MushroomExpert.Com |accessdate=2012-06-15}} 8. ^1 {{cite journal |author1=Largent, David L. |author2=Sugihata, Neil |author3=Brinitzer, Ann |title=Amanita gemmata, a non-host-specific mycorrhizal fungus of Arctostaphylos manzanita |journal=Mycologia |year=1980 |volume=72 |issue=2 |pages=435–439 |jstor=3759273 |url=http://www.cybertruffle.org.uk/cyberliber/59350/0072/002/0435.htm | doi = 10.2307/3759273 }} 9. ^1 {{cite book |author1=McKnight, Vera B. |author2=McKnight, Kent H. |title=A Field Guide to Mushrooms: North America | series=Peterson Field Guides |publisher=Houghton Mifflin |location=Boston, Massachusetts |year=1987 |pages=225–226 |isbn=978-0-395-91090-0 |url=}} 10. ^1 {{cite journal |author1=Drehmel, Dennis |author2=Moncalvo, Jean-Marc |author3=Vilgalys, Rytas |title=Molecular phylogeny of Amanita based on large subunit ribosomal DNA sequences: implications for taxonomy and character evolution |journal=Mycologia |year=1999 |volume=91 |issue=4 |pages=610–618 |doi=10.2307/3761246 |url=http://www.cybertruffle.org.uk/cyberliber/59350/0091/004/0610.htm|jstor=3761246 }} 11. ^1 2 {{cite journal |author1=Moncalvo, Jean-Marc |author2=Drehmel, Dennis |author3=Vilgalys, Rytas |title=Variation in modes and rates of evolution in nuclear and mitochondrial ribosomal DNA in the mushroom genus Amanita (Agaricales, Basidiomycota): phylogenetic implications |journal=Molecular Phylogenetics and Evolution |year=2000 |volume=16 |issue=1 |pages=48–63 |doi=10.1006/mpev.2000.0782 |pmid=10877939}} 12. ^1 {{cite journal |author =Murrill, William A. |title=The Amanitas of Eastern North America |journal=Mycologia |year=1913 |volume=5 |issue=2 |pages=72–86 |doi=10.2307/3753566 |jstor=3753566 }} 13. ^1 {{cite web |author=Russell, Alice B. |title=Amanita gemmata |url=http://www.ces.ncsu.edu/depts/hort/consumer/poison/Amanige.htm |publisher=North Carolina State University |work=Poisonous Plants of North Carolina |year=1997 |accessdate=2012-06-20 |archive-url=https://web.archive.org/web/20131003030020/http://www.ces.ncsu.edu/depts/hort/consumer/poison/Amanige.htm |archive-date=2013-10-03 |dead-url=yes |df= }} 14. ^1 {{cite journal |author1=Nieto, Montserrat Pestaña |author2=Carbone, Serena Santolamazza |title=Characterization of juvenile maritime pine (Pinus pinaster Ait.) ectomycorrhizal fungal community using morphotyping, direct sequencing and fruitbodies sampling |journal=Mycorrhiza |year=2008 |volume=19 |issue=2 |pmid=18972139 |pages=91–98 |doi=10.1007/s00572-008-0207-0}} 15. ^1 {{cite journal |author =Saber, M. |title=The species of Amanita in Iran |journal=Iranian Journal of Plant Pathology |year=1995 |volume=31 |issue=1–4 |pages=15–18 |language=Arabic |issn=0006-2774}} 16. ^1 {{cite journal |author1=Sierralta, A. |author2=Jeria, M.E. |author3=Figueroa, G. |author4=Pinto, J. |author5=Araya, J.C. |author6=San Juan, J. |author7=Grinbergs, J. |author8=Valenzuela, E. |title=Intoxicación por callampas venenosas en la IX Región. Rol de Amanita gemmata |trans-title=Mushroom Poisoning in the 9th region of Chile. Role of Amanita gemmata |journal=Revista Medica de Chile |year=1994 |volume=122 |issue=7 |pages=795–802 |issn=0034-9887 |language=Spanish |pmid=7732230}} 17. ^1 {{cite journal |author1=Tulloss, Rodham E. |author2=Lindgren, Janet E. |title=Amanita aprica—a new toxic species from western North America |journal=Mycotaxon |year=2005 |volume=91 |pages=193–205 |url=http://www.cybertruffle.org.uk/cyberliber/59575/0091/0193.htm}} 18. ^1 2 3 4 5 6 7 {{cite web |url=http://www.amanitaceae.org/?Amanita%20gemmata |title=Amanita gemmata |author =Tulloss, Rodham E. |year=2010 |work=Amanitaceae.org – Taxonomy and Morphology of Amanita and Limacella |accessdate=2012-06-21}} 19. ^1 {{cite web |url=http://www.amanitaceae.org/?stirps%20Gemmata |title=stirps Gemmata |author1=Tulloss, Rodham E. |author2=Yang, Zhu L. |year=2012 |work=Amanitaceae.org – Taxonomy and Morphology of Amanita and Limacella |accessdate=2012-06-21}} 20. ^1 {{cite web |author =Tulloss, Rodham E |title=Amanita orientigemmata |url=http://www.amanitaceae.org/?Amanita%20orientigemmata |publisher=Amanitaceae.org |accessdate=2012-10-26}} 21. ^1 {{cite web |title=Amanita amici Gillet :t.2, 1891 |url=http://www.mycobank.org/BioloMICS.aspx?Table=Mycobank&Rec=235530&Fields=All |publisher=MycoBank. International Mycological Association |accessdate=2012-11-25}} 22. ^1 {{cite web |url=http://www.speciesfungorum.org/Names/SynSpecies.asp?RecordID=202929 |title=Amanita gemmata (Fr.) Bertill. |publisher=Species Fungorum. CAB International |accessdate=2010-12-05}} 23. ^1 2 {{cite web |url=http://www.mycobank.org/MycoTaxo.aspx?Link=T&Rec=202929 |title=Amanita gemmata (Fr.) Bertill. 1866 |publisher=MycoBank. International Mycological Association |accessdate=2010-12-05}} 24. ^1 {{cite journal |author1=Villanueva-Jimenez, Emmanuel |author2=Villegas-Rios, Margarita |author3=Cifuentes-Blanco, Joaquin |author4=Leon-Avendano, Hugo |title=Diversity of the genus Amanita in two areas with different forestry management in Ixtlán de Juárez, Oaxaca, Mexico |journal=Revista Mexicana de Biodiversidad |year=2006 |volume=77 |issue=1 |pages=17–22 |language=Spanish |issn=1870-3453}} 25. ^1 {{cite book |author1=Zhishu, Bi |author2=Zheng, Guoyang |author3=Taihui, Li |title=The Macrofungus Flora of China's Guangdong Province (Chinese University Press) |publisher=Columbia University Press |location=New York, New York |year=1993 |page=273 |isbn=978-962-201-556-2 |url=https://books.google.com/books?id=0cAered-vqYC&pg=PA273}} }} External links
7 : Amanita|Fungi of Asia|Fungi of Europe|Poisonous fungi|Psychoactive fungi|Fungi described in 1838|Taxa named by Elias Magnus Fries |
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