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词条 Centrosaurinae
释义

  1. Classification

  2. Biogeography

  3. Body Size

  4. Reproduction

  5. See also

  6. Footnotes

  7. References

{{Automatic taxobox
| name = Centrosaurines
| taxon = Centrosaurinae
| fossil_range = Late Cretaceous, {{fossil_range|80.8|66}}
| image = Paläontologisches Museum in München Monoclonius.JPG
| image_caption = Centrosaurus "nasicornus" skeleton, Palaeontological Museum Munich
| authority = Lambe, 1915
| type_species = Centrosaurus apertus
| type_species_authority = Lambe, 1904
| subdivision_ranks = Subgroups
| subdivision =
  • Albertaceratops
  • Brachyceratops?
  • Diabloceratops
  • Machairoceratops
  • Monoclonius
  • Medusaceratops
  • Sinoceratops
  • Wendiceratops
  • Xenoceratops
  • Nasutoceratopsini
    • Avaceratops
    • Crittendenceratops
    • Nasutoceratops
    • Yehuecauhceratops
  • Eucentrosaura
    • Centrosaurini
    • Centrosaurus
    • Coronosaurus
    • Rubeosaurus
    • Spinops
    • Styracosaurus
    • Pachyrhinosaurini

| synonyms =
  • Pachyrhinosaurinae Sternberg, 1950
  • Monocloniinae Nopcsa, 1923

}}Centrosaurinae (Greek: pointed lizards) is a subfamily of ceratopsid dinosaurs, a group of large quadrupedal ornithiscians. Centrosaurinae was named by paleontologist Lawrence Lambe in 1915, with Centrosaurus as the type genus. The centrosaurines are further divided into three tribes: the Nasutoceratopsini, the Centrosaurini, and the Pachyrhinosaurini by Ryan et al (2016).[1] Nasutoceratopsins are defined as centrosaurines closer to Nasutoceratops titusi than to Centrosaurus apertus and centrosaurins are defined as centrosaurines (more specifically eucentrosaurans) closer to Centrosaurus apertus than to Pachyrhinosaurus canadensis. The only division used up until then was Pachyrhinosaurini which was defined as centrosaurines closer to Pachyrhinosaurus canadensis than to Centrosaurus apertus. Centrosaurine fossil remains are known primarily from the northern region of Laramidia (modern day Alberta, Montana, and Alaska) but isolated taxa have been found in China and Utah as well.[2] Defining features of centrosaurines include a large nasal horn, short supratemporal horns, and an ornamented frill projecting from the back of the skull.[3] With the exception of Centrosaurus apertus, all adult centrosaurines have spike-like ornaments midway up the skull.[4] Morphometric analysis shows that centrosaurines differ from other ceratopsian groups in skull, snout, and frill shapes.[5] There is evidence to suggest that male centrosaurines had an extended period of adolescence and sexual ornamention did not appear until adulthood.[3]

Classification

The cladogram presented here follows a 2016 phylogenetic analysis by Rivera-Sylva et al..[6]

{{clade| style=font-size:85%; line-height:85%
|label1=Centrosaurinae
|1={{clade
|1=Diabloceratops eatoni
|2={{clade
|label1 =Nasutoceratopsini
|1={{clade
|1=Avaceratops lammersi
|2=Nasutoceratops titusi
|3=Yehuecauhceratops mudei }}
|2={{clade
|1=Xenoceratops foremostensis
|2={{clade
|1={{clade
|1=Sinoceratops zhuchengensis
|2=Wendiceratops pinhornensis }}
|2=Albertaceratops nesmoi
|3={{clade
|label1 =Centrosaurini
|1={{clade
|1=Rubeosaurus ovatus
|2=Styracosaurus albertensis
|3=Coronosaurus brinkmani
|4={{clade
|2=Spinops sternbergorum
|1=Centrosaurus apertus }} }}
|label2 =Pachyrhinosaurini
|2={{clade
|1=Einiosaurus procurvicornis
|2={{clade
|1=Achelousaurus horneri
|2={{clade
|1=Pachyrhinosaurus canadensis
|2=Pachyrhinosaurus lakustai
|3=Pachyrhinosaurus perotorum }} }} }} }} }} }} }} }} }}

The cladogram presented below follows a phylogenetic analysis by Chiba et al. (2017):[7]

{{clade| style=font-size:85%; line-height:85%
|label1=Centrosaurinae
|1={{clade
|1={{clade
|1=Diabloceratops eatoni
|2=Machairoceratops cronusi }}
|2={{clade
|label1 =Nasutoceratopsini
|1={{clade
|1=Avaceratops lammersi (ANSP 15800)
|2=MOR 692
|3=CMN 8804
|4=Nasutoceratops titusi
|5=Malta new taxon }}
|2={{clade
|1=Xenoceratops foremostensis
|2={{clade
|1={{clade
|1=Sinoceratops zhuchengensis
|2=Wendiceratops pinhornensis }}
|2=Albertaceratops nesmoi
|3=Medusaceratops lokii
|label4 =Eucentrosaura
|4={{clade
|label1 =Centrosaurini
|1={{clade
|1={{clade
|1=Rubeosaurus ovatus
|2=Styracosaurus albertensis }}
|2={{clade
|1=Coronosaurus brinkmani
|2={{clade
|2=Spinops sternbergorum
|1=Centrosaurus apertus }} }} }}
|label2 =Pachyrhinosaurini
|2={{clade
|1=Einiosaurus procurvicornis
|label2 =Pachyrostra
|2={{clade
|1=Achelousaurus horneri
|2={{clade
|1=Pachyrhinosaurus canadensis
|2={{clade
|1=Pachyrhinosaurus lakustai
|2=Pachyrhinosaurus perotorum }} }} }} }} }} }} }} }} }} }}

The cladogram presented here represents the conclusions of Dalman et al. (2018).[8]

{{clade| style=font-size:85%; line-height:85%
|1={{clade
|1=Diabloceratops eatoni
|label2=Centrosaurinae
|2={{clade
|label1=Nasutoceratopsini
|1={{clade
|1={{clade
|1={{clade
|1={{clade
|1=Yehuecauhceratops mudei
|2=Nasutoceratops titusi }}
|2={{clade
|1=Avaceratops lammersi
|2=MOR 692 }} }}
|2=CMN 8804 }}
|2={{clade
|1=Crittendenceratops krzyzanowskii
|2='Malta centrosaurine' }} }}
|2={{clade
|1=Albertaceratops nesmoi
|2={{clade
|1=Medusaceratops lokii
|2={{clade
|1={{clade
|1=Sinoceratops zhuchengensis
|2=Wendiceratops pinhornensis }}
|2={{clade
|1=Machairoceratops cronusi
|label2=Eucentrosaura
|2={{clade
|1={{clade
|1={{clade
|1=Einiosaurus procurvicornis
|2=Achelousaurus horneri }}
|2={{clade
|1=Pachyrhinosaurus perotorum
|2={{clade
|1=Pachyrhinosaurus lakustai
|2=Pachyrhinosaurus canadensis }} }} }}
|label2=Centrosaurini
|2={{clade
|1={{clade
|1=Coronosaurus brinkmani
|2={{clade
|1=Xenoceratops foremostensis
|2=Spinops sternbergorum }} }}
|2={{clade
|1=Centrosaurus apertus
|2={{clade
|1=Rubeosaurus ovatus
|2=Styracosaurus albertensis }} }} }} }} }} }} }} }} }} }} }}

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BarData=

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 bar:era          from: 1860    till: 1900    color:1800s    text:19th from: 1900    till: 2000    color:1900s    text:20th from: 2000    till: 2040    color:2000s    text:21st

Biogeography

Centrosaurine fossils have mostly been found in Western North America (Alberta, Montana, & Alaska).[2] In the United States, two taxa, Diabloceratops and Machairoceratops, have been found as far south as Utah. Yehuecauhceratops is a nasutoceratopsin from Coahuila, Mexico and the southernmost occurrence of a centrosaurine in North America. [2] No centrosaurine fossils had been uncovered outside Western North America until 2010 when Sinoceratops zhuchengensis was discovered in the Shandong Province of China.[12] Some authors question the placement of Sinoceratops within Centrosaurinae, however. All other Late Cretaceous dinosaur groups from North America have been found in Asia as well so the initial absence of Asian centrosaurines had been surprising.[12] The current evidence suggests that Centrosaurinae originated in Laramidia 90-80 million years ago.[2] This means Sinoceratops would have migrated to China from North America.[1] Some hypothesize that centrosaurines originated in the south Laramidia and only later radiated north.[9]

Body Size

Compared to their sister group, Chasmosaurinae, centrosaurines are relatively small. The primitive Sinoceratops zhuchengensis is an exception, with an estimated skull length of 180 cm.[10] By contrast, the skull length of Albertoceratops was more typical for this group at only 67 cm.[4] In general, centrosaurines were about the size of a rhinoceros with body lengths ranging from 2.5 to 8 meters.[11]

Reproduction

Possible neonate sized centrosaurine fossils have been documented in the scientific literature. Research indicates that centrosaurines did not achieve fully developed mating signals until nearly fully grown.[3] Scott D. Sampson finds commonality between the slow growth of mating signals in centrosaurines and the extended adolescence of animals whose social structures are ranked hierarchies founded on age-related differences. In these sorts of groups young males are typically sexually mature for several years before actually beginning to breed, when their mating signals are most fully developed. Females, by contrast do not have such an extended adolescence.

See also

{{Portal|Dinosaurs}}
  • Timeline of ceratopsian research

Footnotes

1. ^{{Cite journal|last=Ryan|first=Michael J.|last2=Holmes|first2=Robert|last3=Mallon|first3=Jordan|last4=Loewen|first4=Mark|last5=Evans|first5=David C.|date=2016-10-27|title=A basal ceratopsid (Centrosaurinae: Nasutoceratopsini) from the Oldman Formation (Campanian) of Alberta, Canada|journal=Canadian Journal of Earth Sciences|volume=54|issue=1|pages=1–14|doi=10.1139/cjes-2016-0110|issn=0008-4077}}
2. ^{{Cite journal|last=Sampson|first=Scott D.|last2=Lund|first2=Eric K.|last3=Loewen|first3=Mark A.|last4=Farke|first4=Andrew A.|last5=Clayton|first5=Katherine E.|date=2013-09-07|title=A remarkable short-snouted horned dinosaur from the Late Cretaceous (late Campanian) of southern Laramidia|url=http://rspb.royalsocietypublishing.org/content/280/1766/20131186|journal=Proc. R. Soc. B|language=en|volume=280|issue=1766|pages=20131186|doi=10.1098/rspb.2013.1186|issn=0962-8452|pmc=3730592|pmid=23864598}}
3. ^{{Cite journal|last=Sampson|first=Scott D.|last2=Ryan|first2=Michael J.|last3=Tanke|first3=Darren H.|date=1997-11-01|title=Craniofacial ontogeny in centrosaurine dinosaurs (Ornithischia: Ceratopsidae): taxonomic and behavioral implications|url=https://academic.oup.com/zoolinnean/article/121/3/293/2633685/Craniofacial-ontogeny-in-centrosaurine-dinosaurs|journal=Zoological Journal of the Linnean Society|volume=121|issue=3|pages=293–337|doi=10.1111/j.1096-3642.1997.tb00340.x|issn=0024-4082}}
4. ^{{Cite journal|last=Ryan|first=Michael J.|date=2007-03-01|title=A new basal centrosaurine ceratopsid from the oldman formation, southeastern alberta|journal=Journal of Paleontology|volume=81|issue=2|pages=376–396|doi=10.1666/0022-3360(2007)81[376:ANBCCF]2.0.CO;2|issn=0022-3360}}
5. ^{{Cite journal|last=Maiorino|first=Leonardo|last2=Farke|first2=Andrew A|last3=Kotsakis|first3=Tassos|last4=Piras|first4=Paolo|date=2017|title=Macroevolutionary patterns in cranial and lower jaw shape of ceratopsian dinosaurs (Dinosauria, Ornithischia): phylogeny, morphological integration, and evolutionary rates|url=http://www.evolutionary-ecology.com/issues/v18/n02/ccar3008.pdf|journal=Evolutionary Ecology Research|volume=18|pages=123–167|via=}}
6. ^{{cite journal | last1 = Rivera-Sylva | first1 = H.E. | last2 = Hendrick | first2 = B.P. | last3 = Dodson | first3 = P. | title = A Centrosaurine (Dinosauria: Ceratopsia) from the Aguja Formation (Late Campanian) of Northern Coahuila, Mexico | journal = PLoS ONE | date = 2016 | volume = 11 | issue = 4 | page = e0150529 | doi = 10.1371/journal.pone.0150529 | pmid = 27073969 | pmc = 4830452 }}
7. ^{{Cite journal|author1=Kentaro Chiba |author2=Michael J. Ryan |author3=Federico Fanti |author4=Mark A. Loewen |author5=David C. Evans |year=2018 |title=New material and systematic re-evaluation of Medusaceratops lokii (Dinosauria, Ceratopsidae) from the Judith River Formation (Campanian, Montana) |journal=Journal of Paleontology |volume=in press |issue= 2|pages= 272–288|doi=10.1017/jpa.2017.62 }}
8. ^{{cite journal|first1=Sebastian G. |last1=Dalman |first2=John-Paul M. |last2=Hodnett |first3=Asher J. |last3=Lichtig |first4=Spencer G. |last4=Lucas |year=2018 |title=A new ceratopsid dinosaur (Centrosaurinae: Nasutoceratopsini) from the Fort Crittenden Formation, Upper Cretaceous (Campanian) of Arizona |journal=New Mexico Museum of Natural History and Science Bulletin |volume=79 |pages=141–164 |url=https://www.researchgate.net/publication/328637301}}
9. ^{{Cite journal|last=Lund|first=Eric K.|last2=O’Connor|first2=Patrick M.|last3=Loewen|first3=Mark A.|last4=Jinnah|first4=Zubair A.|date=2016-05-18|title=A New Centrosaurine Ceratopsid, Machairoceratops cronusi gen et sp. nov., from the Upper Sand Member of the Wahweap Formation (Middle Campanian), Southern Utah|journal=PLOS ONE|volume=11|issue=5|pages=e0154403|doi=10.1371/journal.pone.0154403|issn=1932-6203|pmc=4871575|pmid=27192148}}
10. ^{{Cite journal|last=Xu|first=Xing|last2=Wang|first2=KeBai|last3=Zhao|first3=XiJin|last4=Li|first4=DunJing|date=2010-06-01|title=First ceratopsid dinosaur from China and its biogeographical implications|journal=Chinese Science Bulletin|language=en|volume=55|issue=16|pages=1631–1635|doi=10.1007/s11434-009-3614-5|issn=1001-6538}}
11. ^{{Cite book|title=Dinosaurs : the most complete, up-to-date encyclopedia for dinosaur lovers of all ages|last=V.|first=Rey, Luis|date=2007|publisher=Random House|isbn=9780375824197|oclc=930042495}}

References

  • {{cite journal | last1 = Sampson | first1 = S. D. | year = 1995b | title = Two new horned dinosaurs from the Upper Cretaceous Two Medicine Formation of Montana; with a phylogenetic analysis of the Centrosaurinae (Ornithischia: Ceratopsidae) | url = | journal = Journal of Vertebrate Paleontology | volume = 15 | issue = 4| pages = 743–760 | doi=10.1080/02724634.1995.10011259}}
  • Sampson, S. D., 2001, Speculations on the socioecology of Ceratopsid dinosaurs (Orinthischia: Neoceratopsia): In: Mesozoic Vertebrate Life, edited by Tanke, D. H., and Carpenter, K., Indiana University Press, pp. 263–276.
  • Tanke, D.H. and Brett-Surman, M.K. 2001. Evidence of Hatchling and Nestling-Size Hadrosaurs (Reptilia:Ornithischia) from Dinosaur Provincial Park (Dinosaur Park Formation: Campanian), Alberta, Canada. pp. 206–218. In: Mesozoic Vertebrate Life—New Research Inspired by the Paleontology of Philip J. Currie. Edited by D.H. Tanke and K. Carpenter. Indiana University Press: Bloomington. xviii + 577 pp.
{{Marginocephalia|T.}}{{Taxonbar|from=Q132304}}

2 : Ceratopsids|Centrosaurines

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