词条 | Centrosaurinae |
释义 |
| name = Centrosaurines | taxon = Centrosaurinae | fossil_range = Late Cretaceous, {{fossil_range|80.8|66}} | image = Paläontologisches Museum in München Monoclonius.JPG | image_caption = Centrosaurus "nasicornus" skeleton, Palaeontological Museum Munich | authority = Lambe, 1915 | type_species = Centrosaurus apertus | type_species_authority = Lambe, 1904 | subdivision_ranks = Subgroups | subdivision =
| synonyms =
}}Centrosaurinae (Greek: pointed lizards) is a subfamily of ceratopsid dinosaurs, a group of large quadrupedal ornithiscians. Centrosaurinae was named by paleontologist Lawrence Lambe in 1915, with Centrosaurus as the type genus. The centrosaurines are further divided into three tribes: the Nasutoceratopsini, the Centrosaurini, and the Pachyrhinosaurini by Ryan et al (2016).[1] Nasutoceratopsins are defined as centrosaurines closer to Nasutoceratops titusi than to Centrosaurus apertus and centrosaurins are defined as centrosaurines (more specifically eucentrosaurans) closer to Centrosaurus apertus than to Pachyrhinosaurus canadensis. The only division used up until then was Pachyrhinosaurini which was defined as centrosaurines closer to Pachyrhinosaurus canadensis than to Centrosaurus apertus. Centrosaurine fossil remains are known primarily from the northern region of Laramidia (modern day Alberta, Montana, and Alaska) but isolated taxa have been found in China and Utah as well.[2] Defining features of centrosaurines include a large nasal horn, short supratemporal horns, and an ornamented frill projecting from the back of the skull.[3] With the exception of Centrosaurus apertus, all adult centrosaurines have spike-like ornaments midway up the skull.[4] Morphometric analysis shows that centrosaurines differ from other ceratopsian groups in skull, snout, and frill shapes.[5] There is evidence to suggest that male centrosaurines had an extended period of adolescence and sexual ornamention did not appear until adulthood.[3] ClassificationThe cladogram presented here follows a 2016 phylogenetic analysis by Rivera-Sylva et al..[6] {{clade| style=font-size:85%; line-height:85%|label1=Centrosaurinae |1={{clade |1=Diabloceratops eatoni |2={{clade |label1 =Nasutoceratopsini |1={{clade |1=Avaceratops lammersi |2=Nasutoceratops titusi |3=Yehuecauhceratops mudei }} |2={{clade |1=Xenoceratops foremostensis |2={{clade |1={{clade |1=Sinoceratops zhuchengensis |2=Wendiceratops pinhornensis }} |2=Albertaceratops nesmoi |3={{clade |label1 =Centrosaurini |1={{clade |1=Rubeosaurus ovatus |2=Styracosaurus albertensis |3=Coronosaurus brinkmani |4={{clade |2=Spinops sternbergorum |1=Centrosaurus apertus }} }} |label2 =Pachyrhinosaurini |2={{clade |1=Einiosaurus procurvicornis |2={{clade |1=Achelousaurus horneri |2={{clade |1=Pachyrhinosaurus canadensis |2=Pachyrhinosaurus lakustai |3=Pachyrhinosaurus perotorum }} }} }} }} }} }} }} }} }} The cladogram presented below follows a phylogenetic analysis by Chiba et al. (2017):[7] {{clade| style=font-size:85%; line-height:85%|label1=Centrosaurinae |1={{clade |1={{clade |1=Diabloceratops eatoni |2=Machairoceratops cronusi }} |2={{clade |label1 =Nasutoceratopsini |1={{clade |1=Avaceratops lammersi (ANSP 15800) |2=MOR 692 |3=CMN 8804 |4=Nasutoceratops titusi |5=Malta new taxon }} |2={{clade |1=Xenoceratops foremostensis |2={{clade |1={{clade |1=Sinoceratops zhuchengensis |2=Wendiceratops pinhornensis }} |2=Albertaceratops nesmoi |3=Medusaceratops lokii |label4 =Eucentrosaura |4={{clade |label1 =Centrosaurini |1={{clade |1={{clade |1=Rubeosaurus ovatus |2=Styracosaurus albertensis }} |2={{clade |1=Coronosaurus brinkmani |2={{clade |2=Spinops sternbergorum |1=Centrosaurus apertus }} }} }} |label2 =Pachyrhinosaurini |2={{clade |1=Einiosaurus procurvicornis |label2 =Pachyrostra |2={{clade |1=Achelousaurus horneri |2={{clade |1=Pachyrhinosaurus canadensis |2={{clade |1=Pachyrhinosaurus lakustai |2=Pachyrhinosaurus perotorum }} }} }} }} }} }} }} }} }} }} The cladogram presented here represents the conclusions of Dalman et al. (2018).[8] {{clade| style=font-size:85%; line-height:85%|1={{clade |1=Diabloceratops eatoni |label2=Centrosaurinae |2={{clade |label1=Nasutoceratopsini |1={{clade |1={{clade |1={{clade |1={{clade |1=Yehuecauhceratops mudei |2=Nasutoceratops titusi }} |2={{clade |1=Avaceratops lammersi |2=MOR 692 }} }} |2=CMN 8804 }} |2={{clade |1=Crittendenceratops krzyzanowskii |2='Malta centrosaurine' }} }} |2={{clade |1=Albertaceratops nesmoi |2={{clade |1=Medusaceratops lokii |2={{clade |1={{clade |1=Sinoceratops zhuchengensis |2=Wendiceratops pinhornensis }} |2={{clade |1=Machairoceratops cronusi |label2=Eucentrosaura |2={{clade |1={{clade |1={{clade |1=Einiosaurus procurvicornis |2=Achelousaurus horneri }} |2={{clade |1=Pachyrhinosaurus perotorum |2={{clade |1=Pachyrhinosaurus lakustai |2=Pachyrhinosaurus canadensis }} }} }} |label2=Centrosaurini |2={{clade |1={{clade |1=Coronosaurus brinkmani |2={{clade |1=Xenoceratops foremostensis |2=Spinops sternbergorum }} }} |2={{clade |1=Centrosaurus apertus |2={{clade |1=Rubeosaurus ovatus |2=Styracosaurus albertensis }} }} }} }} }} }} }} }} }} }} }} ImageSize = width:1000px height:auto barincrement:15px PlotArea = left:10px bottom:50px top:10px right:10px Period = from:1860 till:2040 TimeAxis = orientation:horizontal ScaleMajor = unit:year increment:50 start:1860 ScaleMinor = unit:year increment:10 start:1860 TimeAxis = orientation:hor AlignBars = justify Colors = #legends id:CAR value:claret id:ANK value:rgb(0.4,0.3,0.196) id:HER value:teal id:HAD value:green id:OMN value:blue id:black value:black id:white value:white id:1900s value:rgb(0.94,0.25,0.24) id:2000s value:rgb(0.2,0.7,0.79) id:2000syears value:rgb(0.52,0.81,0.91) id:1900syears value:rgb(0.95,0.56,0.45) id:1700s value:rgb(0.5,0.78,0.31) id:1700syears value:rgb(0.63,0.78,0.65) id:latecretaceous value:rgb(0.74,0.82,0.37) id:1800syears value:rgb(0.95,0.98,0.11) id:paleogene value:rgb(0.99,0.6,0.32) id:paleocene value:rgb(0.99,0.65,0.37) id:eocene value:rgb(0.99,0.71,0.42) id:oligocene value:rgb(0.99,0.75,0.48) id:1800s value:rgb(0.999999,0.9,0.1) id:miocene value:rgb(0.999999,0.999999,0) id:pliocene value:rgb(0.97,0.98,0.68) id:quaternary value:rgb(0.98,0.98,0.5) id:pleistocene value:rgb(0.999999,0.95,0.68) id:holocene value:rgb(0.999,0.95,0.88) BarData= bar:eratop bar:space bar:periodtop bar:space bar:NAM1 bar:NAM2 bar:NAM3 bar:NAM4 bar:NAM5 bar:NAM6 bar:NAM7 bar:NAM8 bar:NAM9 bar:NAM10 bar:NAM11 bar:NAM12 bar:NAM13 bar:NAM14 bar:NAM15 bar:NAM16 bar:NAM17 bar:space bar:period bar:space bar:era PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 shift:(7,-4) bar:periodtop from: 1860 till: 1870 color:1800syears text:60s from: 1870 till: 1880 color:1800syears text:70s from: 1880 till: 1890 color:1800syears text:80s from: 1890 till: 1900 color:1800syears text:90s from: 1900 till: 1910 color:1900syears text:00s from: 1910 till: 1920 color:1900syears text:10s from: 1920 till: 1930 color:1900syears text:20s from: 1930 till: 1940 color:1900syears text:30s from: 1940 till: 1950 color:1900syears text:40s from: 1950 till: 1960 color:1900syears text:50s from: 1960 till: 1970 color:1900syears text:60s from: 1970 till: 1980 color:1900syears text:70s from: 1980 till: 1990 color:1900syears text:80s from: 1990 till: 2000 color:1900syears text:90s from: 2000 till: 2010 color:2000syears text:00s from: 2010 till: 2020 color:2000syears text:10s from: 2020 till: 2030 color:2000syears text:20s from: 2030 till: 2040 color:2000syears text:30s bar:eratop from: 1860 till: 1900 color:1800s text:19th from: 1900 till: 2000 color:1900s text:20th from: 2000 till: 2040 color:2000s text:21st PlotData= color:1900s bar:NAM1 at:1904 mark:(line,black) text:Centrosaurus color:1900s bar:NAM1 at:1904 mark:(line,black) text:Centrosaurus color:1800s bar:NAM1 at:1876 mark:(line,black) text:Monoclonius color:1900s bar:NAM2 at:1913 mark:(line,black) text:Styracosaurus color:1900s bar:NAM1 at:1986 mark:(line,black) text:Avaceratops color:1900s bar:NAM1 at:1950 mark:(line,black) text:Pachyrhinosaurus color:1900s bar:NAM2 at:1995 mark:(line,black) text:Achelousaurus color:1800s bar:NAM3 at:1995 mark:(line,black) text:Einiosaurus color:1900s bar:NAM5 at:2007 mark:(line,black) text:Albertaceratops color:1900s bar:NAM6 at:2010 mark:(line,black) text:Diabloceratops color:1900s bar:NAM7 at:2010 mark:(line,black) text:Medusaceratops color:1900s bar:NAM8 at:2010 mark:(line,black) text:Rubeosaurus color:1900s bar:NAM9 at:2010 mark:(line,black) text:Sinoceratops color:1900s bar:NAM10 at:2011 mark:(line,black) text:Spinops color:1800s bar:NAM11 at:2012 mark:(line,black) text:Coronosaurus color:1800s bar:NAM12 at:2012 mark:(line,black) text:Xenoceratops color:1900s bar:NAM1 at:2013 mark:(line,black) text:Nasutoceratops color:1900s bar:NAM2 at:2015 mark:(line,black) text:Wendiceratops color:1900s bar:NAM3 at:2016 mark:(line,black) text:Machairoceratops color:1900s bar:NAM4 at:2017 mark:(line,black) text:Yehuecauhceratops color:1900s bar:NAM13 at:2018 mark:(line,black) text:Crittendenceratops PlotData= bar:period from: 1860 till: 1870 color:1800syears text:60s from: 1870 till: 1880 color:1800syears text:70s from: 1880 till: 1890 color:1800syears text:80s from: 1890 till: 1900 color:1800syears text:90s from: 1900 till: 1910 color:1900syears text:00s from: 1910 till: 1920 color:1900syears text:10s from: 1920 till: 1930 color:1900syears text:20s from: 1930 till: 1940 color:1900syears text:30s from: 1940 till: 1950 color:1900syears text:40s from: 1950 till: 1960 color:1900syears text:50s from: 1960 till: 1970 color:1900syears text:60s from: 1970 till: 1980 color:1900syears text:70s from: 1980 till: 1990 color:1900syears text:80s from: 1990 till: 2000 color:1900syears text:90s from: 2000 till: 2010 color:2000syears text:00s from: 2010 till: 2020 color:2000syears text:10s from: 2020 till: 2030 color:2000syears text:20s from: 2030 till: 2040 color:2000syears text:30s bar:era from: 1860 till: 1900 color:1800s text:19th from: 1900 till: 2000 color:1900s text:20th from: 2000 till: 2040 color:2000s text:21st BiogeographyCentrosaurine fossils have mostly been found in Western North America (Alberta, Montana, & Alaska).[2] In the United States, two taxa, Diabloceratops and Machairoceratops, have been found as far south as Utah. Yehuecauhceratops is a nasutoceratopsin from Coahuila, Mexico and the southernmost occurrence of a centrosaurine in North America. [2] No centrosaurine fossils had been uncovered outside Western North America until 2010 when Sinoceratops zhuchengensis was discovered in the Shandong Province of China.[12] Some authors question the placement of Sinoceratops within Centrosaurinae, however. All other Late Cretaceous dinosaur groups from North America have been found in Asia as well so the initial absence of Asian centrosaurines had been surprising.[12] The current evidence suggests that Centrosaurinae originated in Laramidia 90-80 million years ago.[2] This means Sinoceratops would have migrated to China from North America.[1] Some hypothesize that centrosaurines originated in the south Laramidia and only later radiated north.[9] Body SizeCompared to their sister group, Chasmosaurinae, centrosaurines are relatively small. The primitive Sinoceratops zhuchengensis is an exception, with an estimated skull length of 180 cm.[10] By contrast, the skull length of Albertoceratops was more typical for this group at only 67 cm.[4] In general, centrosaurines were about the size of a rhinoceros with body lengths ranging from 2.5 to 8 meters.[11] ReproductionPossible neonate sized centrosaurine fossils have been documented in the scientific literature. Research indicates that centrosaurines did not achieve fully developed mating signals until nearly fully grown.[3] Scott D. Sampson finds commonality between the slow growth of mating signals in centrosaurines and the extended adolescence of animals whose social structures are ranked hierarchies founded on age-related differences. In these sorts of groups young males are typically sexually mature for several years before actually beginning to breed, when their mating signals are most fully developed. Females, by contrast do not have such an extended adolescence. See also{{Portal|Dinosaurs}}
Footnotes1. ^1 {{Cite journal|last=Ryan|first=Michael J.|last2=Holmes|first2=Robert|last3=Mallon|first3=Jordan|last4=Loewen|first4=Mark|last5=Evans|first5=David C.|date=2016-10-27|title=A basal ceratopsid (Centrosaurinae: Nasutoceratopsini) from the Oldman Formation (Campanian) of Alberta, Canada|journal=Canadian Journal of Earth Sciences|volume=54|issue=1|pages=1–14|doi=10.1139/cjes-2016-0110|issn=0008-4077}} 2. ^1 2 3 {{Cite journal|last=Sampson|first=Scott D.|last2=Lund|first2=Eric K.|last3=Loewen|first3=Mark A.|last4=Farke|first4=Andrew A.|last5=Clayton|first5=Katherine E.|date=2013-09-07|title=A remarkable short-snouted horned dinosaur from the Late Cretaceous (late Campanian) of southern Laramidia|url=http://rspb.royalsocietypublishing.org/content/280/1766/20131186|journal=Proc. R. Soc. B|language=en|volume=280|issue=1766|pages=20131186|doi=10.1098/rspb.2013.1186|issn=0962-8452|pmc=3730592|pmid=23864598}} 3. ^1 2 {{Cite journal|last=Sampson|first=Scott D.|last2=Ryan|first2=Michael J.|last3=Tanke|first3=Darren H.|date=1997-11-01|title=Craniofacial ontogeny in centrosaurine dinosaurs (Ornithischia: Ceratopsidae): taxonomic and behavioral implications|url=https://academic.oup.com/zoolinnean/article/121/3/293/2633685/Craniofacial-ontogeny-in-centrosaurine-dinosaurs|journal=Zoological Journal of the Linnean Society|volume=121|issue=3|pages=293–337|doi=10.1111/j.1096-3642.1997.tb00340.x|issn=0024-4082}} 4. ^1 {{Cite journal|last=Ryan|first=Michael J.|date=2007-03-01|title=A new basal centrosaurine ceratopsid from the oldman formation, southeastern alberta|journal=Journal of Paleontology|volume=81|issue=2|pages=376–396|doi=10.1666/0022-3360(2007)81[376:ANBCCF]2.0.CO;2|issn=0022-3360}} 5. ^{{Cite journal|last=Maiorino|first=Leonardo|last2=Farke|first2=Andrew A|last3=Kotsakis|first3=Tassos|last4=Piras|first4=Paolo|date=2017|title=Macroevolutionary patterns in cranial and lower jaw shape of ceratopsian dinosaurs (Dinosauria, Ornithischia): phylogeny, morphological integration, and evolutionary rates|url=http://www.evolutionary-ecology.com/issues/v18/n02/ccar3008.pdf|journal=Evolutionary Ecology Research|volume=18|pages=123–167|via=}} 6. ^{{cite journal | last1 = Rivera-Sylva | first1 = H.E. | last2 = Hendrick | first2 = B.P. | last3 = Dodson | first3 = P. | title = A Centrosaurine (Dinosauria: Ceratopsia) from the Aguja Formation (Late Campanian) of Northern Coahuila, Mexico | journal = PLoS ONE | date = 2016 | volume = 11 | issue = 4 | page = e0150529 | doi = 10.1371/journal.pone.0150529 | pmid = 27073969 | pmc = 4830452 }} 7. ^{{Cite journal|author1=Kentaro Chiba |author2=Michael J. Ryan |author3=Federico Fanti |author4=Mark A. Loewen |author5=David C. Evans |year=2018 |title=New material and systematic re-evaluation of Medusaceratops lokii (Dinosauria, Ceratopsidae) from the Judith River Formation (Campanian, Montana) |journal=Journal of Paleontology |volume=in press |issue= 2|pages= 272–288|doi=10.1017/jpa.2017.62 }} 8. ^{{cite journal|first1=Sebastian G. |last1=Dalman |first2=John-Paul M. |last2=Hodnett |first3=Asher J. |last3=Lichtig |first4=Spencer G. |last4=Lucas |year=2018 |title=A new ceratopsid dinosaur (Centrosaurinae: Nasutoceratopsini) from the Fort Crittenden Formation, Upper Cretaceous (Campanian) of Arizona |journal=New Mexico Museum of Natural History and Science Bulletin |volume=79 |pages=141–164 |url=https://www.researchgate.net/publication/328637301}} 9. ^{{Cite journal|last=Lund|first=Eric K.|last2=O’Connor|first2=Patrick M.|last3=Loewen|first3=Mark A.|last4=Jinnah|first4=Zubair A.|date=2016-05-18|title=A New Centrosaurine Ceratopsid, Machairoceratops cronusi gen et sp. nov., from the Upper Sand Member of the Wahweap Formation (Middle Campanian), Southern Utah|journal=PLOS ONE|volume=11|issue=5|pages=e0154403|doi=10.1371/journal.pone.0154403|issn=1932-6203|pmc=4871575|pmid=27192148}} 10. ^1 2 {{Cite journal|last=Xu|first=Xing|last2=Wang|first2=KeBai|last3=Zhao|first3=XiJin|last4=Li|first4=DunJing|date=2010-06-01|title=First ceratopsid dinosaur from China and its biogeographical implications|journal=Chinese Science Bulletin|language=en|volume=55|issue=16|pages=1631–1635|doi=10.1007/s11434-009-3614-5|issn=1001-6538}} 11. ^{{Cite book|title=Dinosaurs : the most complete, up-to-date encyclopedia for dinosaur lovers of all ages|last=V.|first=Rey, Luis|date=2007|publisher=Random House|isbn=9780375824197|oclc=930042495}} References
2 : Ceratopsids|Centrosaurines |
随便看 |
|
开放百科全书收录14589846条英语、德语、日语等多语种百科知识,基本涵盖了大多数领域的百科知识,是一部内容自由、开放的电子版国际百科全书。