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词条 Caprinae
释义

  1. Characteristics

  2. Evolution

  3. Extant species

  4. Fossil genera

  5. References

{{hatnote|Not to be confused with Antilocapridae. "Goat-antelope" redirects here. For the species sometimes called goat antelope see, Tibetan antelope.}}{{Automatic taxobox
| fossil_range = {{fossil range|Late Miocene|present}}
| image = Stone Sheep British Columbia.jpg
| image_caption = Stone sheep (Ovis dalli stonei) in British Columbia, 2009
| taxon = Caprinae
| authority = Gray, 1821
| subdivision_ranks = Genera
| subdivision =

  Nemorhaedus

  Rupicapra

  Oreamnos

  Budorcas

  Ovibos

  Hemitragus

  Ammotragus

  Pseudois

  Capra

  Ovis

and see text


}}

The subfamily Caprinae is part of the ruminant family Bovidae, and consists of mostly medium-sized bovids. A member of this subfamily is called a caprine.{{citation needed|date=March 2019|reason=According to Wiktionary, a caprine is a member of the tribe Caprini. A member of the subfamily Caprinae would be called a caprid.}} A member is also sometimes referred to as a goat-antelope, however, this term "goat-antelope" does not mean that these animals are true antelopes: a true antelope is a bovid with a cervid-like or antilocaprid-like morphology.

Within this subfamily Caprinae, a prominent tribe, Caprini, includes sheep and goats.

Some earlier taxonomies considered Caprinae a separate family called Capridae (whence a caprid), but now it is usually considered a subfamily within the family Bovidae, whence a caprine is a kind of bovid.

Characteristics

Although most goat-antelopes are gregarious and have fairly stocky builds, they diverge in many other ways – the muskox (Ovibos moschatus) is adapted to the extreme cold of the tundra; the Rocky Mountain goat (Oreamnos americanus) of North America is specialised for very rugged terrain; the urial (Ovis orientalis) occupies a largely infertile area from Kashmir to Iran, including much desert country. The European mouflon (Ovis musimon) is thought to be the ancestor of the modern domestic sheep (Ovis aries).

Many species have become extinct since the last ice age, probably largely because of human interaction. Of the survivors:

  • Five are classified as endangered,
  • Eight as vulnerable,
  • Seven as of concern and needing conservation measures, but at lower risk, and
  • Seven species are secure.

Members of the group vary considerably in size, from just over {{convert|1|m|ft|sigfig=1|abbr=on}} long for a full-grown grey goral (Nemorhaedus goral), to almost {{convert|2.5|m|sigfig=1|abbr=on}} long for a musk ox, and from under {{convert|30|kg|abbr=on}} to more than {{convert|350|kg|abbr=on}}. Musk oxen in captivity have reached over {{convert|650|kg|abbr=on}}.

The lifestyles of caprids fall into two broad classes: 'resource-defenders', which are territorial and defend a small, food-rich area against other members of the same species; and 'grazers', which gather together into herds and roam freely over a larger, usually relatively infertile area.

The resource-defenders are the more primitive group: they tend to be smaller in size, dark in colour, males and females fairly alike, have long, tassellated ears, long manes, and dagger-shaped horns. The grazers (sometimes collectively known as tsoan caprids, from the Hebrew tso'n meaning sheep and goats) evolved more recently. They tend to be larger, highly social, and rather than mark territory with scent glands, they have highly evolved dominance behaviours. No sharp line divides the groups, but a continuum varies from the serows at one end of the spectrum to sheep, true goats, and musk oxen at the other.

Evolution

The goat-antelope, or caprid, group is known from as early as the Miocene, when members of the group resembled the modern serow in their general body form.[1] The group did not reach its greatest diversity until the recent ice ages, when many of its members became specialised for marginal, often extreme, environments: mountains, deserts, and the subarctic region.

The ancestors of the modern sheep and goats (both rather vague and ill-defined terms) are thought to have moved into mountainous regions – sheep becoming specialised occupants of the foothills and nearby plains, and relying on flight and flocking for defence against predators, and goats adapting to very steep terrain where predators are at a disadvantage.

Extant species

{{update-section|date=August 2017}}FAMILY BOVIDAE
  • Subfamily Caprinae
    • Tribe Ovibovini{{contradict-inline|article=muskox|date=May 2015}}
    • Genus Budorcas
    • Takin, Budorcas taxicolor
    • Genus Ovibos
    • Muskox, Ovibos moschatus
    • {{Anchor|Tribe Caprini}}Tribe Caprini
    • Genus Ammotragus
    • Barbary sheep, Ammotragus lervia
    • Genus Arabitragus
    • Arabian tahr, Arabitragus jayakari
    • Genus Capra
    • West Caucasian tur, Capra caucasica
    • East Caucasian tur, Capra caucasica cylindricornis
    • Markhor, Capra falconeri
    • Wild goat, Capra aegagrus
    • Domestic goat, Capra aegagrus hircus
    • Alpine ibex, Capra ibex
    • Nubian ibex, Capra nubiana
    • Spanish ibex, Capra pyrenaica
    • Siberian ibex, Capra sibirica
    • Walia ibex, Capra walie
    • Genus Hemitragus
    • Himalayan tahr, Hemitragus jemlahicus
    • Genus Ovis
    • Argali, Ovis ammon
    • Domestic sheep, Ovis aries
    • Bighorn sheep, Ovis canadensis
    • Dall or thinhorn sheep, Ovis dalli
    • European mouflon, Ovis musimon
    • Marco Polo sheep, Ovis polii
    • Snow sheep, Ovis nivicola
    • Wild sheep, Ovis orientalis
    • Mouflon, Ovis orientalis orientalis
    • Urial, Ovis orientalis vignei
    • Genus Nilgiritragus
    • Nilgiri tahr, Nilgiritragus hylocrius
    • Genus Pseudois
    • Bharal (Himalayan blue sheep), Pseudois nayaur
    • Dwarf blue sheep, Pseudois schaeferi
    • Tribe Naemorhedini
    • Genus Capricornis
    • Japanese serow, Capricornis crispus
    • Sumatran serow, Capricornis sumatraensis
    • Taiwan serow, Capricornis swinhoei
    • Chinese serow, Capricornis milneedwardsii
    • Red serow, Capricornis rubidus
    • Himalayan serow Capricornis thar
    • Genus Nemorhaedus
    • Red goral, Nemorhaedus baileyi
    • Chinese goral, Nemorhaedus griseus
    • Grey goral, Nemorhaedus goral
    • Long-tailed goral, Naemorhedus caudatus
    • Genus Oreamnos
    • Mountain goat, Oreamnos americanus
    • Genus Rupicapra
    • Pyrenean chamois, Rupicapra pyrenaica
    • Alpine chamois, Rupicapra rupicapra

Fossil genera

The following extinct genera of Caprinae have been identified:[2][3]

  • Tribe Ovibovini
    • Genus Bootherium
    • Bootherium bombifrons
    • Genus Euceratherium
    • Euceratherium collinum
    • Genus Makapania
    • Makapania broomi
    • Genus Soergelia
    • Soergelia mayfieldi
    • Genus Tsaidamotherium
    • Tsaidamotherium brevirostrum
    • Tsaidamotherium hedini
  • {{Anchor|Tribe Caprini}}Tribe Caprini
    • Genus Myotragus
    • Myotragus balearicus
Unsorted{{div col|colwidth=15em}}
  • Benicerus
  • Boopsis
  • Capraoryx
  • Caprotragoides
  • Criotherium
  • Damalavus
  • Gallogoral
  • Lyrocerus
  • Megalovis
  • Mesembriacerus
  • Neotragocerus
  • Nesogoral
  • Norbertia
  • Numidocapra
  • Oioceros
  • Olonbulukia
  • Pachygazella
  • Pachytragus
  • Palaeoreas
  • Palaeoryx
  • Paraprotoryx
  • Parapseudotragus
  • Parurmiatherium
  • Praeovibos
  • Procamptoceras
  • Prosinotragus
  • Protoryx
  • Pseudotragus
  • Samotragus
  • Sinocapra
  • Sinomegoceros
  • Sinopalaeoceros
  • Sinotragus
  • Sivacapra
  • Sporadotragus
  • Symbos
  • Tethytragus
  • Tossunnoria
  • Turcocerus
  • Urmiatherium
{{div col end}}

References

{{more citations needed|date=October 2015}}
1. ^{{Cite book|editor=Macdonald, D.|author= Geist, Valerius|year=1984 |title= The Encyclopedia of Mammals|publisher= Facts on File|location=New York|pages= 584–587|isbn= 0-87196-871-1}}
2. ^tolweb.org
3. ^palaeos.org
{{Artiodactyla|R.3}}{{Taxonbar|from=Q189804}}

4 : Caprids|Extant Miocene first appearances|Taxa named by John Edward Gray|Mammal subfamilies

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