词条 | Methanosarcinaceae |
释义 |
|image=Metba.gif |image_caption=Methanosarcina barkeri fusaro | domain = Archaea | regnum = Euryarchaeota | phylum = Euryarchaeota | classis = Methanomicrobia | ordo = Methanosarcinales | familia = Methanosarcinaceae | subdivision_ranks = Genera | subdivision =
| binomial = Methanosarcinaceae | binomial_authority = Balch and Wolfe 1981 }} In taxonomy, the Methanosarcinaceae are a family of the Methanosarcinales.[1] PhylogenyThe currently accepted taxonomy is based on the List of Prokaryotic names with Standing in Nomenclature (LPSN)[2] and National Center for Biotechnology Information (NCBI)[3] and the phylogeny is based on 16S rRNA-based LTP release 106 by 'The All-Species Living Tree' Project.[4] {{Clade|1={{clade |1=?Halomethanococcus doii ♦ Yu and Kawamura 1988 |2=?Methanohalobium evestigatum Zhilina and Zavarzin 1988 |3=Methanosalsum zhilinae (Mathrani et al. 1988) Boone and Baker 2002 |4={{clade |label1=Methanomethylovorans |1={{clade |1=?M. victoriae ♠ Muyodi et al. 2000 |2=M. hollandica Lomans et al. 2004 (type sp.) |3=M. thermophila Jiang et al. 2005 |2={{clade |1={{clade |1=Methanimicrococcus blatticola Sprenger et al. 2000 |label2=Methanosarcina |2={{clade |1=?M. horonobensis Shimizu et al. 2011 |2=?M. methanica (Smit 1930) Kluyver and van Niel 1936 (nom. rejic.) |3=?M. siciliae (Stetter and König 1989) Ni et al. 1994 emend. Elberson and Sowers 1997 |4=?M. vacuolata Zhilina and Zavarzin 1987 |5={{clade |1=M. semesiae Lyimo et al. 2000 |2=M. baltica von Klein et al. 2002 emend. Singh et al. 2005 |6={{clade |1=M. lacustris Simankova et al. 2002 |2={{clade |1=M. mazei (Barker 1936) Mah and Kuhn 1984 |2={{clade |1=M. acetivorans Sowers et al. 1986 |2={{clade |1=M. barkeri Schnellen 1947 emend. Maestrojuán et al. 1992 (type sp.) |2=M. thermophila Zinder et al. 1985 }} }} }} }} }} }} |2={{clade |1={{clade |label1=Methanococcoides |1={{clade |1=?M. methylutens Sowers and Ferry 1985 (type sp.) |2=M. alaskense Singh et al. 2005 |3=M. burtonii Franzmann et al. 1993 |label2=Methanohalophilus |2={{clade |1=?M. euhalobius ♠ (Obraztsova et al. 1987) Davidova et al. 1997 |2=?M. halophilus (Zhilina 1984) Wilharm et al. 1991 |3=M. mahii Paterek and Smith 1988 (type sp.) |4=M. portucalensis Boone et al. 1993 }} }} |label2=Methanolobus |2={{clade |1=?M. bombayensis Kadam et al. 1994 |2=?M. psychrophilus ♠ Zhang et al. 2008 |3=?M. tindarius König and Stetter 1983 (type sp.) |4=M. zinderi Doerfert et al. 2009 |5={{clade |1=M. oregonensis (Liu et al. 1990) Boone 2002 |2={{clade |1=M. taylorii Oremland and Boone 1994 |2={{clade |1=M. profundi Mochimaru et al. 2009 |2=M. vulcani Stetter et al. 1989 emend. Kadam and Boone 1995 }} }} }} }} }} }} }} }} }} Notes: ♠ Strains found at the National Center for Biotechnology Information (NCBI) but not listed in the List of Prokaryotic names with Standing in Nomenclature (LPSN) ♦ Type strain lost or not available BiochemistryA notable trait of Methanosarcinaceae is that they are methanogens that incorporate the unusual amino acid pyrrolysine into their enzymes.[5] The enzyme monomethylamine methyltransferase catalyzes the reaction of monomethylamine to methane. This enzyme includes pyrrolysine. The unusual amino acid is inserted using a unique tRNA, the anticodon of which is UAG. In most organisms, and in most Methanosarcinaceae proteins, UAG is a stop codon. However in this enzyme, and anywhere else pyrrolysine is incorporated, likely through contextual markers on the mRNA, the pyrrolysine-loaded tRNA is inserted instead of the release factor. They also have a unique aminoacyl-tRNA synthetase to specifically load this tRNA with pyrrolysine. This unique adaptation is still the subject of significant study. References1. ^See the NCBI [https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=2206 webpage on Methanosarcinaceae]. Data extracted from the {{cite web | url=ftp://ftp.ncbi.nih.gov/pub/taxonomy/ | title=NCBI taxonomy resources | publisher=National Center for Biotechnology Information | accessdate=2007-03-19}} 2. ^{{cite web|author=J.P. Euzéby |url=http://www.bacterio.cict.fr/classifphyla.html#Euryarchaeota |title=Methanomicrobia |accessdate=2011-11-17 |publisher=List of Prokaryotic names with Standing in Nomenclature (LPSN) |deadurl=yes |archiveurl=https://web.archive.org/web/20130127030659/http://www.bacterio.cict.fr/classifphyla.html |archivedate=2013-01-27 |df= }} 3. ^{{cite web |author = Sayers|display-authors = etal| url=https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Tree&id=224756&lvl=5&lin |title=Methanomicrobia |accessdate=2011-06-05 |publisher=National Center for Biotechnology Information (NCBI) taxonomy database}} 4. ^'The All-Species Living Tree' Project.{{cite web | url=http://www.arb-silva.de/fileadmin/silva_databases/living_tree/LTP_release_106/LTPs106_SSU_tree.pdf | title=16S rRNA-based LTP release 106 (full tree) | publisher=Silva Comprehensive Ribosomal RNA Database | accessdate=2011-11-17}} 5. ^Lehninger A, Nelson D, Cox M. Lehninger principles of biochemistry. 6th ed. New York: W.H. Freeman; 2013 p. 1124-1126. Further readingScientific journals
Scientific books
Scientific databases{{Taxonomic references|taxon=Methanosarcinaceae}}External links{{Taxonomic links|microbe=yes|NCBI_taxID=2206|taxoname=Methanosarcinaceae|LSPN_letter=m|LSPN_taxoname=methanosarcinaceae}}{{Archaea classification}}{{Taxonbar|from=Q1949357}} 2 : Archaea taxonomic families|Euryarchaeota |
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