| 词条 | Tylopilus |
| 释义 |
| image = Tylopilus felleus 060914c.jpg | image_width = 240px | image_caption = Tylopilus felleus | regnum = Fungi | divisio = Basidiomycota | classis = Agaricomycetes | ordo = Boletales | familia = Boletaceae | genus = Tylopilus | genus_authority = P.Karst. (1881) | type_species = Tylopilus felleus | type_species_authority = (Bull.) P.Karst. (1881) | synonyms_ref = [1] | synonyms =
}} Tylopilus is a genus of over 100 species of mycorrhizal bolete fungi separated from Boletus. Its best known member is the bitter bolete (Tylopilus felleus), the only species found in Europe. More species are found in North America, such as the edible species T. alboater, and the tropics. Australia is another continent where many species are found. All members of the genus form mycorrhizal relationships with trees. Members of the genus are distinguished by their pinkish pore surfaces. TaxonomyThe genus was first defined by Petter Adolf Karsten in 1881.[6] The type species, Tylopilus felleus, was originally described in 1788 as a species of Boletus by French mycologist Pierre Bulliard.[7] Tylopilus means "bumpy or swollen pileus",[8] from the Greek tylos "bump" and pilos "hat".[9] Molecular analysis indicates the genus, like other large genera within the Boletales, is polyphyletic.[10] A lineage of Tylopilus chromapes (now Harrya chromapes and related species) has been shown to be only distantly related to other members of Tylopilus. Hence T. chromapes is now the type species of the new genus Harrya and, related to it, several Australian species moved to Australopilus.[11] T. valens was also moved to its own genus, Pseudoaustroboletus.[12]DescriptionFruit bodies of the genus Tylopilus are encountered as large stout bolete mushrooms, which generally arise from the ground or occasionally from wood. They have stout stipes, which do not have a ring.[8] A key field character which distinguishes them from members of the genus Boletus is the presence of their pink-tinged pores (though these may be white when young).[14] The "pink pore" feature is a polyphyletic morphology that does not unite the Tylopilus species using traditional morphological characters (Smith and Thiers or Singer's concepts). The spore print manifests various shades of pinkish-brown, through reddish-brown and even chocolate brown.[8] EdibilityMany species have a bitter taste and are inedible,[8] a key feature in identification.[14] The black velvet bolete (T. alboater) is edible and tasty, but often ignored.[8] Species{{As of|2015|August}}, Index Fungorum lists 111 valid species of Tylopilus.[19] About 40 are found in western North America.[8] A large number have been recorded from Australia, with 26 aligned with existing taxa and another 15 not assignable. Members of the genus are also abundant in South America, particularly in forests with trees of the genus Dicymbe in Guyana, as well as Central America and elsewhere across tropical regions around the world.[21] All are mycorrhizal.{{Div col}}
References1. ^1 {{cite journal |author=Beck G. |title=Versuch einer systematischen Gliederung der Gattung Boletus L. em. |journal=Zeitschrift für Pilzkunde |year=1923 |volume=2 |issue=7 |pages=141–49 |language=German}} [1][2][3][4][5][6][7][8][9][10][11][12][13][14][15][16]2. ^1 2 3 4 5 6 {{cite book |vauthors=Bessette AR, Bessette A, Roody WC |title=North American Boletes: A Color Guide to the Fleshy Pored Mushrooms |publisher=Syracuse University Press |location=Syracuse, New York |year=2000 |page=256 |isbn=0-8156-0588-9 |url=https://books.google.com/books?id=geiwy6Jf_ZcC&pg=PA256}} 3. ^1 {{cite journal |vauthors=Binder M, Hibbett DS |year=2006 |title=Molecular systematics and biological diversification of Boletales |volume=98 |issue=6 |pages=971–81 |url=http://www.clarku.edu/faculty/dhibbett/Reprints%20PDFs/Binder%20and%20Hibbett%202006%20Boletales.pdf |format=PDF |deadurl=yes |archiveurl=https://web.archive.org/web/20111226013340/http://www.clarku.edu/faculty/dhibbett/Reprints%20PDFs/Binder%20and%20Hibbett%202006%20Boletales.pdf |archivedate=2011-12-26 |df= }} 4. ^1 {{cite book |author=Bulliard JBF |title=Herbier de la France |year=1788 |volume=8 |at=plate 379 |publisher=Chez l'auteur, Didot, Debure, Belin |location=Paris |language=French |url=https://www.biodiversitylibrary.org/page/4309231}} 5. ^1 {{cite web |author=Kirk PM |title=Species Fungorum (version 26th August 2015). In: Species 2000 & ITIS Catalogue of Life |url=http://www.catalogueoflife.org/col/browse/tree/id/aef3fa4f00c544da6802d194f7a621b3 |accessdate=2015-08-29}} 6. ^1 {{cite journal |vauthors=Halling RE, Nuhn M, Osmundson T, Fechner N, Trappe JM, Soytong K, Arora D, Hibbett DS, Binder M |title=Affinities of the Boletus chromapes group to Royoungia and the description of two new genera, Harrya and Australopilus |journal=Australian Systematic Botany |year=2012 |volume=25 |pages=418–31 |doi=10.1071/SB12028}} 7. ^1 {{cite journal |author=Karsten PA |title=Enumeratio Boletinearum et Polyporearum Fennicarum, systemate novo dispositarum |journal=Revue mycologique, Toulouse |year=1881 |volume=3 |issue=9 |pages=16–19 |url=https://www.biodiversitylibrary.org/page/11599311}} 8. ^1 2 {{cite web |url=http://www.mushroomexpert.com/tylopilus.html |title=The genus Tylopilus |last=Kuo |first=Michael |date=March 2005 |accessdate=23 August 2011}} 9. ^1 {{cite journal |vauthors=((Li Y-C)), Li F, ((Zeng N-K)), ((Cui Y-Y)), Yang ZL |title=A new genus Pseudoaustroboletus (Boletaceae, Boletales) from Asia as inferred from molecular and morphological data |journal=Mycological Progress |volume=13 |year=2014 |doi=10.1007/s11557-014-1011-1 |id=1011}} 10. ^1 {{cite journal |author=Murrill WA |title=The Boletaceae of North America – 1 |journal=Mycologia |year=1909 |volume=1 |issue=1 |pages=4–18 (see p. 15) |url=http://www.cybertruffle.org.uk/cyberliber/59350/0001/001/0017.htm |doi=10.2307/3753167}} 11. ^1 {{cite book |vauthors=Nilson S, Persson O |year=1977 |title = Fungi of Northern Europe 1: Larger Fungi (Excluding Gill-Fungi) |publisher=Penguin |location=Harmondsworth, England |isbn=0-14-063005-8 |pages=102–03}} 12. ^1 {{cite journal |author=Snell WH |title=New proposals relating to the genera of the Boletaceae |journal=Mycologia |year=1942 |volume=34 |issue=4 |pages=403–11 |url=http://www.cybertruffle.org.uk/cyberliber/59350/0034/004/0403.htm |doi=10.2307/3754982}} 13. ^1 {{cite journal |author=Takahashi H. |title=Two new species and one new combination of Agaricales from Japan |journal=Mycoscience |year=2002 |volume=43 |issue=5 |pages=397–403 |doi=10.1007/s102670200058}} 14. ^1 {{cite journal |author=Takahashi H. |title=Five new species of the Boletaceae from Japan |journal=Mycoscience |year=2007 |volume=48 |issue=2 |pages=90–9 |doi=10.1007/s10267-006-0332-6p}} 15. ^1 {{cite web |title=Tylopilus P. Karst. 1881 |url=http://www.mycobank.org/MycoTaxo.aspx?Link=T&Rec=18700 |publisher=MycoBank. International Mycological Association |accessdate=30 Aug 2011}} 16. ^1 {{cite journal |author=Watling R. |year=2001 |title=Australian Boletes: Their Diversity and Possible Origins|journal=Australian Systematic Botany |volume=14 |issue=3 |pages=407–16 |doi=10.1071/SB99031}} }} External links
3 : Tylopilus|Boletales genera|Taxa named by Petter Adolf Karsten |
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