| 词条 | Maranhão red-handed howler |
| 释义 |
| name = Maranhão red-handed howler | status = EN | status_system = IUCN3.1 | status_ref = [1] | genus = Alouatta | species = ululata | authority = Elliot, 1912 | synonyms = Alouatta belzebul ululata | range_map = Maranhao Red-handed Howler area.png | range_map_caption = Maranhao Red-handed Howler range }} The Maranhão red-handed howler (Alouatta ululata) is an endangered species of howler monkey endemic to forests (for example Babaçu forests) in the northeastern Brazilian states of Ceará, Maranhão and Piauí.[1] It was previously thought to be a subspecies of the red-handed howler, but unlike that species, the Maranhão red-handed howler is strongly sexually dichromatic.[2] References1. ^1 {{Cite journal | author = de Oliveira, M.M. | author2 = Kierulff, M.C.M. | last-author-amp = yes | title = Alouatta ululata | journal = The IUCN Red List of Threatened Species | volume = 2008 | page = e.T918A13094890 | publisher = IUCN | date = 2008 | url = http://www.iucnredlist.org/details/918/0 | doi = 10.2305/IUCN.UK.2008.RLTS.T918A13094890.en | access-date = 26 December 2017}} {{Atelidae nav}}{{Taxonbar|from=Q1353729}}{{DEFAULTSORT:howler, red-handed, Maranhão}}{{newworld-monkey-stub}}2. ^Gregorin, R. (2006). {{Pt icon}} Taxonomia e variação geográfica das espécies do gênero Alouatta Lacépède (Primates, Atelidae) no Brasil. Rev. Bras. Zool. 23(1). 4 : Howler monkeys of South America|Mammals of Brazil|Endemic fauna of Brazil|Mammals described in 1912 |
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