请输入您要查询的百科知识：

　

词条 Neanderthal extinction
释义

  1. Coexistence prior to extinction

  2. Possible cause of extinction

     Violence  Parasites and pathogens  Competitive replacement   Species specific disadvantages   Division of labor  Anatomical differences and running ability  Modern humans' advantage in hunting warm climate animals  Interbreeding  Climate change  Natural catastrophe 

  3. See also

  4. References

  5. External links

{{short description|Causes and mechanism of the extinction of the Neanderthal people}}Neanderthal extinction began around 40,000 years ago in the Paleolithic Europe, after anatomically modern humans had reached the continent. This date, which is based on research published in Nature in 2014, is much earlier than previous estimates, and it was established through improved radio carbon dating methods analyzing 40 sites from Spain to Russia. The survey did not include sites in Asia, where Neanderthals may have survived longer.[1] Evidence for continued Neanderthal presence in the Iberian Peninsula at 37,000 years ago was published in 2017.[1]

Hypotheses on the fate of the Neanderthals include violence from encroaching anatomically modern humans,[2] parasites and pathogens, competitive replacement, competitive exclusion, extinction by interbreeding with early modern human populations,[3] natural catastrophes, and failure or inability to adapt to climate change. It is unlikely that any one of these hypotheses is sufficient on its own; rather, multiple factors probably contributed to the demise of an already widely-dispersed population.[4][5]

Coexistence prior to extinction

In research published in Nature in 2014, an analysis of radiocarbon dates from forty Neanderthal sites from Spain to Russia found that the Neanderthals disappeared in Europe between 41,000 and 39,000 years ago with 95% probability. The study also found with the same probability that modern humans and Neanderthals overlapped in Europe for between 2,600 and 5,400 years.[6] Modern humans reached Europe between 45,000 and 43,000 years ago.[7] Improved radiocarbon dating published in 2015 indicates that Neanderthals disappeared around 40,000 years ago, which overturns older carbon dating which indicated that Neanderthals may have lived as recently as 24,000 years ago,[8] including in refugia on the south coast of the Iberian peninsula such as Gorham's Cave.[9]

Zilhão et al. (2017) argue for pushing this date forward by some 3,000 years, to 37,000 years ago.[1]

Inter-stratification of Neanderthal and modern human remains has been suggested,[10] but is disputed.[11]

Possible cause of extinction

Violence

Some authors have discussed the possibility that Neanderthal extinction was either precipitated or hastened by violent conflict with Homo sapiens. Conflict and warfare are virtually ubiquitous features of hunter-gatherer societies, including conflicts over limited resources, such as prey and water.[12] It is therefore plausible to suggest that violence, including primitive warfare, would have transpired between the two human species.[13] The hypothesis that early humans violently replaced Neanderthals was first proposed by French palaeontologist Marcellin Boule (the first person to publish an analysis of a Neanderthal) in 1912.[14] Several finds in both Homo-sapiens and Neanderthal bones indicate inter-species aggression from injuries (grooves in the bones themselves) that could only have come from spear or other projectile tips crafted with prevalent tool-making methods contemporary to the time.[15]{{verify credibility|date=May 2018}}

Parasites and pathogens

Another possibility is the spread among the Neanderthal population of pathogens or parasites carried by Homo sapiens.[16] Neanderthals would have limited immunity to diseases they had not been exposed to, so diseases carried into Europe by Homo sapiens could have been particularly lethal to them if Homo sapiens were relatively resistant. If it were relatively easy for pathogens to leap between these two similar species, perhaps because they lived in close proximity, then Homo sapiens would have provided a pool of individuals capable of infecting Neanderthals and potentially preventing the epidemic from burning itself out as Neanderthal population fell. On the other hand, the same mechanism could work in reverse, and the resistance of Homo sapiens to Neanderthal pathogens and parasites would need explanation. An examination of human and Neanderthal genomes and adaptations regarding pathogens or parasites may shed light on this issue.

Competitive replacement

Species specific disadvantages

Slight competitive advantage on the part of modern humans has accounted for Neanderthals' decline on a timescale of thousands of years.[17]

Generally small and widely-dispersed fossil sites suggest that Neanderthals lived in less numerous and socially more isolated groups than contemporary Homo sapiens. Tools such as Mousterian flint stone flakes and Levallois points are remarkably sophisticated from the outset, yet they have a slow rate of variability and general technological inertia is noticeable during the entire fossil period. Artifacts are of utilitarian nature, and symbolic behavioral traits are undocumented before the arrival of modern humans in Europe around 40,000 to 35,000 years ago.[18][19]

Jared Diamond, supporter of competitive replacement, points out in his book The Third Chimpanzee that the genocidal replacement of Neanderthals by modern humans is comparable to patterns of behavior that occur whenever people with advanced technology clash with less advanced people.[20]

Division of labor

In 2006, two anthropologists of the University of Arizona proposed an efficiency explanation for the demise of the Neanderthals.[21] In an article titled "What's a Mother to Do? The Division of Labor among Neanderthals and Modern Humans in Eurasia",[22] it was posited that Neanderthal division of labor between the sexes was less developed than Middle paleolithic Homo sapiens. Both male and female Neanderthals participated in the single occupation of hunting big game, such as bison, deer, gazelles and wild horses. This hypothesis proposes that the Neanderthal's relative lack of labor division resulted in less efficient extraction of resources from the environment as compared to Homo sapiens.

Anatomical differences and running ability

Researchers such as Karen L. Steudel of the University of Wisconsin have highlighted the relationship of Neanderthal anatomy (shorter and stockier than that of modern humans{{citation needed|date=March 2013}}) and the ability to run and the requirement of energy (30% more).[23]{{Failed verification|date=January 2011}}

Nevertheless, in the recent study, researchers Martin Hora and Vladimir Sladek of Charles University in Prague show that Neanderthal lower limb configuration, particularly the combination of robust knees, long heels and short lower limbs, increased the effective mechanical advantage of Neanderthal knee and ankle extensors, thus reducing the force needed and the energy spent for locomotion significantly. The walking cost of the Neanderthal male is now estimated to be 8–12% higher than that of anatomically modern males, whereas the walking cost of the Neanderthal female is considered to be virtually equal to that of anatomically modern females.[24]

Other researchers, like Yoel Rak, from Tel-Aviv University in Israel, have noted that the fossil records show that Neanderthal pelvises in comparison to modern human pelvises would have made it much harder for Neanderthals to absorb shock and to bounce off from one step to the next, giving modern humans another advantage over Neanderthals in running and walking ability. However, Rak also notes that all archaic humans had wide pelves, indicating that this is the ancestral morphology and that modern humans experienced a shift towards narrower pelves late in Pleistocene. [25]

Modern humans' advantage in hunting warm climate animals

Pat Shipman, from Pennsylvania State University in the United States, argues that the domestication of the dog gave modern humans an advantage when hunting.[26] The oldest remains of domesticated dogs were found in Belgium (31,700 BP) and in Siberia (33,000 BP).[27][28] A survey of early sites of modern humans and Neanderthals with faunal remains across Spain, Portugal and France provided an overview of what modern humans and Neanderthals ate.[29] Rabbit became more frequent, while large mammals – mainly eaten by the Neanderthals – became increasingly rare. In 2013, DNA testing on the "Altai dog", a paleolithic dog's remains from the Razboinichya Cave (Altai Mountains), has linked this 33,000-year-old dog with the present lineage of Canis lupus familiaris.[30]

Interbreeding

{{main|Archaic human admixture with modern humans}}{{Further|Neanderthal Genome Project}}

Interbreeding can only account for a certain degree of Neanderthal population decrease. A homogeneous absorption of an entire species is a rather unrealistic idea. This would also be counter to strict versions of the Recent African Origin, since it would imply that at least part of the genome of Europeans would descend from Neanderthals, whose ancestors left Africa at least 350,000 years ago.

The most vocal proponent of the hybridization hypothesis is Erik Trinkaus of Washington University.[31][32] Trinkaus claims various fossils as hybrid individuals, including the "child of Lagar Velho", a skeleton found at Lagar Velho in Portugal.[33] In a 2006 publication co-authored by Trinkaus, the fossils found in 1952 in the cave of Peștera Muierilor, Romania, are likewise claimed as hybrids.[34]

Genetic studies indicate some form of hybridization between archaic humans and modern humans had taken place after modern humans emerged from Africa. An estimated 1–4% of the DNA in Europeans and Asians (e.g. French, Chinese and Papua probands) is non-modern, and shared with ancient Neanderthal DNA rather than with sub-Saharan Africans (e.g. Yoruba and San probands).[35]

Modern-human findings in Abrigo do Lagar Velho, Portugal allegedly featuring Neanderthal admixtures have been published.[36] However, the interpretation of the Portuguese specimen is disputed.[37]

Jordan, in his work Neanderthal, points out that without some interbreeding, certain features on some "modern" skulls of Eastern European Cro-Magnon heritage are hard to explain. In another study, researchers have recently found in Peştera Muierilor, Romania, remains of European humans from ~37,000–42,000 years ago[38]who possessed mostly diagnostic "modern" anatomical features, but also had distinct Neanderthal features not present in ancestral modern humans in Africa, including a large bulge at the back of the skull, a more prominent projection around the elbow joint, and a narrow socket at the shoulder joint.

The Neanderthal genome project published papers in 2010 and 2014 stating that Neanderthals contributed to the DNA of modern humans, including most humans outside sub-Saharan Africa, as well as a few populations in sub-Saharan Africa, through interbreeding, likely between 50,000 and 60,000 years ago.[39][40][41] Recent studies also show that a few Neanderthals began mating with ancestors of modern humans long before the large "out of Africa migration" of the present day non-Africans, as early as 100,000 years ago.[42] In 2016, research indicated that there were three distinct episodes of interbreeding between modern humans and Neanderthals: the first encounter involved the ancestors of non-African modern humans, probably soon after leaving Africa; the second, after the ancestral Melanesian group had branched off (and subsequently had a unique episode of interbreeding with Denisovans); and the third, involving the ancestors of East Asians only.[43]

{{wide image|Neanderthal DNA Comparison (SharedDNA) T1a.png|1000px|align-cap=center|Neanderthal DNA Comparison (SharedDNA)}}

While interbreeding is viewed as the most parsimonious interpretation of the genetic discoveries, the authors point out they cannot conclusively rule out an alternative scenario, in which the source population of non-African modern humans was already more closely related to Neanderthals than other Africans were, due to ancient genetic divisions within Africa. Among the genes shown to differ between present-day humans and Neanderthals were RPTN, SPAG17, CAN15, TTF1 and PCD16.

Climate change

{{see also|Neanderthals of Gibraltar#Lifestyle of the Gibraltar Neanderthals}}

Neanderthals went through a demographic crisis in Western Europe that seems to coincide with climate change that resulted in a period of extreme cold in Western Europe. "The fact that Neanderthals in Western Europe were nearly extinct, but then recovered long before they came into contact with modern humans came as a complete surprise to us," said Love Dalén, associate professor at the Swedish Museum of Natural History in Stockholm. If so, this would indicate that Neanderthals may have been very sensitive to climate change.[44]

Natural catastrophe

{{main|Campanian Ignimbrite Eruption}}

A number of researchers have argued that the Campanian Ignimbrite Eruption, a volcanic eruption near Naples, Italy, about 39,280 ± 110 years ago (older estimate ~37,000 years), erupting about {{convert|200|km3|cumi|sigfig=2|abbr=on}} of magma ({{convert|500|km3|cumi|sigfig=2|abbr=on}} bulk volume) contributed to the extinction of Neanderthal man.[45] The argument has been developed by Golovanova et al.[46][47] The hypothesis posits that although Neanderthals had encountered several Interglacials during 250,000 years in Europe,[48] inability to adapt their hunting methods caused their extinction facing H. sapiens competition when Europe changed into a sparsely vegetated steppe and semi-desert during the last Ice Age.[49] Studies of sediment layers at Mezmaiskaya Cave suggest a severe reduction of plant pollen.[47] The damage to plant life would have led to a corresponding decline in plant-eating mammals hunted by the Neanderthals.[47][50][51]

See also

  • {{annotated link|List of human evolution fossils|List of hominina (hominid) fossils}}
  • {{annotated link|Quaternary extinction event}}

References

1. ^1 João Zilhão et al., "Precise dating of the Middle-to-Upper Paleolithic transition in Murcia (Spain) supports late Neandertal persistence in Iberia", Heliyon 3(11), 16 November 2017, {{doi|10.1016/j.heliyon.2017.e00435}}.
2. ^{{cite journal |vauthors=Hortolà P, Martínez-Navarro B |title=The Quaternary megafaunal extinction and the fate of Neanderthals: An integrative working hypothesis|journal=Quaternary International|volume=295|pages=69–72|year=2013 |doi=10.1016/j.quaint.2012.02.037|bibcode = 2013QuInt.295...69H }}
3. ^Interbreeding took place in western Asia between about 65,000 and 47,000 years ago, see {{cite journal| last1=Kuhlwilm|first1=M.| last2=Gronau|first2=I.| last3=Hubisz|first3=M.J.| last4=de Filippo|first4=C.| last5=Prado-Martinez|first5=J.| last6=Kircher|first6=M.| display-authors=etal |title=Ancient gene flow from early modern humans into Eastern Neanderthals| journal=Nature| date=2016| volume=530| issue=7591| pages=429–433| doi=10.1038/nature16544|pmid=26886800|bibcode=2016Natur.530..429K|pmc=4933530}}
4. ^{{cite web|url=http://www.anthropologyinpractice.com/2010/10/what-are-those-darned-neanderthals-up.html |title=What Are Those Darned Neanderthals Up to Now? |publisher=Anthropology in practice |date=October 11, 2010 |accessdate=September 20, 2016}}
5. ^{{cite web|url=http://scienceforwriters.blogspot.com/2015/03/did-volcanic-eruption-kill-off.html |title=Did a Volcanic Eruption Kill Off the Neandertals? |publisher=Science for writers |date=March 25, 2015 |accessdate=September 20, 2016}}
6. ^1 {{cite journal|journal=Nature|last=Higham|first= Tom|display-authors=etal|title=The timing and spatiotemporal patterning of Neanderthal disappearance|date=21 August 2014|doi=10.1038/nature13621|volume=512|issue=7514|pages=306–309|pmid=25143113|bibcode=2014Natur.512..306H}}
7. ^{{cite web|author=John Noble Wilford |url=https://www.nytimes.com/2011/11/03/science/fossil-teeth-put-humans-in-europe-earlier-than-thought.html?scp=1&sq=kents%20cavern&st=cse&_r=0 |title=Fossil Teeth Put Humans in Europe Earlier Than Thought |publisher=Nytimes.com |date=2011-11-02 |accessdate=2017-02-23}}
8. ^{{cite web |last=Dorey |first=Fran |title=Homo Neanderthalensis – The Neanderthals |url=https://australianmuseum.net.au/homo-neanderthalensis |date=30 October 2015 |work=Australian Museum |accessdate=1 April 2017 }}
9. ^{{cite news|url=http://news.bbc.co.uk/1/hi/5343266.stm?lsm|title=Neanderthals' 'last rock refuge'|last=Rincon|first=Paul|date=13 September 2006|work=BBC News|accessdate= 11 October 2009}}{{cite journal |vauthors=Finlayson C, Pacheco FG, Rodríguez-Vidal J, etal |title=Late survival of Neanderthals at the southernmost extreme of Europe |journal=Nature |volume=443 |issue=7113 |pages= 850–853 |date=October 2006 |pmid=16971951 |doi=10.1038/nature05195|bibcode = 2006Natur.443..850F |hdl=10261/18685 }}
10. ^{{cite journal |vauthors=Gravina B, Mellars P, Ramsey CB |title=Radiocarbon dating of interstratified Neanderthal and early modern human occupations at the Chatelperronian type-site |journal=Nature |volume=438 |issue=7064 |pages= 51–56 |date=November 2005 |pmid=16136079 |doi=10.1038/nature04006 |bibcode=2005Natur.438...51G}}
11. ^{{cite journal|last=Zilhão|first=João |author2=Francesco d’Errico |author3=Jean-Guillaume Bordes |author4=Arnaud Lenoble |author5=Jean-Pierre Texier |author6=Jean-Philippe Rigaud|year=2006|title=Analysis of Aurignacian interstratification at the Châtelperronian-type site and implications for the behavioral modernity of Neandertals|journal=PNAS|volume=103|issue=33|pages= 12643–12648|pmid=16894152|doi=10.1073/pnas.0605128103|pmc=1567932|bibcode=2006PNAS..10312643Z}}
12. ^{{cite web|last1=Hawks|first1=John|title=Hunter-gatherer mortality|url=http://johnhawks.net/weblog/reviews/life_history/aging_evolution/hill_2007_hiwi_mortality.html|website=john hawks weblog|accessdate=3 March 2016}}
13. ^{{cite journal|last1=Ko|first1=Kwang Hyun|title=Hominin interbreeding and the evolution of human variation|journal=Journal of Biological Research-Thessaloniki|date=2016|volume=23|pages=17|doi=10.1186/s40709-016-0054-7|pmid=27429943|pmc=4947341}}
14. ^Boule, M 1920, Les hommes fossiles, Masson, Paris.
15. ^{{cite web |url=http://www.livescience.com/7794-human-stabbed-neanderthal-evidence-suggests.html |title=Human Stabbed a Neanderthal, Evidence Suggests |publisher=Livescience.com |date= |accessdate= January 21, 2017}}
16. ^{{cite news|last1=Underdown|first1=Simon|title=Brookes research finds modern humans gave fatal diseases to Neanderthals|url=https://www.brookes.ac.uk/about-brookes/news/brookes-research-finds-modern-humans-gave-fatal-diseases-to-neanderthals/|accessdate=3 March 2016|publisher=Oxford Brookes University news|date=10 April 2015}}
17. ^{{cite journal |last1=Banks |first1=William E. |last2=d'Errico |first2=Francesco |last3=Peterson |first3=A. Townsend |last4=Kageyama |first4=Masa |last5=Sima |first5=Adriana |last6=Sánchez-Goñi |first6=Maria-Fernanda |date=24 December 2008 |editor1-last=Harpending |editor1-first=Henry |title=Neanderthal Extinction by Competitive Exclusion |journal=PLoS ONE |volume=3 |issue=12 |pages=e3972 |issn=1932-6203 |doi=10.1371/journal.pone.0003972 |pmid=19107186 |pmc=2600607 |bibcode=2008PLoSO...3.3972B}}
18. ^{{cite web|url=http://eol.org/pages/4454114/details |title=Homo neanderthalensis Brief Summary |publisher=EOL |date= |accessdate=September 26, 2016}}
19. ^{{cite journal | title =Symbolic or utilitarian? Juggling interpretations of Neanderthal behavior: new inferences from the study of engraved stone surfaces | pmid=25020018 | doi=10.4436/JASS.92007 | volume=92 | issue=92 | journal=J Anthropol Sci | pages=233–55 | last1 = Peresani | first1 = M | last2 = Dallatorre | first2 = S | last3 = Astuti | first3 = P | last4 = Dal Colle | first4 = M | last5 = Ziggiotti | first5 = S | last6 = Peretto | first6 = C| year=2014 }}
20. ^Diamond, J. (1992). The Third Chimpanzee: The Evolution and Future of the Human Animal. New York: Harper Collins, p. 45.
21. ^Nicholas Wade, [https://www.nytimes.com/2006/12/05/science/05nean.html "Neanderthal Women Joined Men in the Hunt"], from The New York Times, 5 December 2006
22. ^{{cite journal | last1 = Kuhn | first1 = Steven L. | last2 = Stiner | first2 = Mary C. | year = 2006| title = What's a Mother to Do? The Division of Labor among Neandertals and Modern Humans in Eurasia | journal = Current Anthropology | volume = 47| issue = 6| pages = 953–981| doi = 10.1086/507197 }}
23. ^{{cite web | first = Karen L. | last = Steudel | title = Karen L. Steudel |publisher=Department of Zoology – University of Wisconsin |location=Madison, Wisconsin | url = http://www.zoology.wisc.edu/faculty/Ste/Ste.html | year = 2009 | accessdate =December 24, 2011}}
24. ^{{cite journal | last1 = Hora | first1 = M | last2 = Sládek | first2 = V | year = 2014 | title = Influence of lower limb configuration on walking cost in Late Pleistocene humans | url = | journal = Journal of Human Evolution | volume = 67| issue = | pages = 19–32| doi = 10.1016/j.jhevol.2013.09.011 | pmid = 24485350 }}
25. ^{{cite web | title = Science: Neanderthals puzzle the anthropologists | publisher = New Scientist | date = 27 April 1991 | first = Roger | last = Lewin | accessdate = January 1, 2011 | url = https://www.newscientist.com/article/mg13017664.500-science-neanderthals-puzzle-the-anthropologists-.html}}
26. ^{{cite journal | last1 = Shipman | first1 = P | year = 2012 | title = Dog domestication may have helped humans thrive while Neandertals declined | url = http://www.americanscientist.org/issues/pub/do-the-eyes-have-it/1 | journal = American Scientist | volume = 100 | issue = 3| page = 198 | doi = 10.1511/2012.96.198 }}
27. ^{{cite journal | last1 = Ovodov | first1 = ND | last2 = Crockford | first2 = SJ | last3 = Kuzmin | first3 = YV | last4 = Higham | first4 = TFG | last5 = Hodgins | first5 = GWL | display-authors = etal | year = 2011 | title = A 33,000-Year-Old Incipient Dog from the Altai Mountains of Siberia: Evidence of the Earliest Domestication Disrupted by the Last Glacial Maximum | url = | journal = PLoS ONE | volume = 6 | issue = 7| page = e22821 | doi = 10.1371/journal.pone.0022821 | pmid=21829526 | pmc=3145761| bibcode = 2011PLoSO...622821O }}
28. ^{{cite journal | last1 = Germonpré | first1 = M. | last2 = Sablin | first2 = M.V. | last3 = Stevens | first3 = R.E. | last4 = Hedges | first4 = R.E.M. | last5 = Hofreiter | first5 = M. | last6 = Stiller | first6 = M. | last7 = Jaenicke-Desprese | first7 = V. | year = 2009 | title = Fossil dogs and wolves from Palaeolithic sites in Belgium, the Ukraine and Russia: osteometry, ancient DNA and stable isotopes | url = | journal = Journal of Archaeological Science | volume = 36 | issue = 2| pages = 473–490 | doi=10.1016/j.jas.2008.09.033}}
29. ^{{cite journal| doi=10.1016/j.jhevol.2013.01.002 | pmid=23422239 | volume=64 | issue=4 | title=Rabbits and hominin survival in Iberia | year=2013 | journal=Journal of Human Evolution | pages=233–241 | last1 = Fa | first1 = John E. | last2 = Stewart | first2 = John R. | last3 = Lloveras | first3 = Lluís | last4 = Vargas | first4 = J. Mario}}
30. ^{{cite journal | last1 = Druzhkova | first1 = AS | last2 = Thalmann | first2 = O | last3 = Trifonov | first3 = VA | last4 = Leonard | first4 = JA | last5 = Vorobieva | first5 = NV | display-authors = etal | year = 2013 | title = Ancient DNA Analysis Affirms the Canid from Altai as a Primitive Dog | url = | journal = PLoS ONE | volume = 8 | issue = 3| page = e57754 | doi = 10.1371/journal.pone.0057754 | pmid=23483925 | pmc=3590291| bibcode = 2013PLoSO...857754D }}
31. ^{{cite journal | last1 = Jones | first1 = Dan | year = 2007 | title = The Neanderthal within | url = | journal = New Scientist | volume = 193 | issue = 2593| pages = 28–32 | doi=10.1016/s0262-4079(07)60550-8}}
32. ^[https://news.yahoo.com/s/hsn/20061030/hl_hsn/modernhumansneanderthalsmayhaveinterbred Modern Humans, Neanderthals May Have Interbred]; Humans and Neanderthals interbred {{webarchive|url=https://web.archive.org/web/20090222210101/http://www.cosmosmagazine.com/node/814 |date=2009-02-22 }}
33. ^{{cite web | title = The Lagar Velho 1 Skeleton | url = http://www.talkorigins.org/faqs/homs/lagarvelho.html | publisher = TalkOrigins | year = 2000 | accessdate = January 1, 2011}}; {{cite news | title = Life on the edge: was a Gibraltar cave last outpost of the lost neanderthal? | first = Ian | last = Sample | publisher = The Guardian | date = 14 September 2006 | accessdate = January 1, 2011 | url = https://www.theguardian.com/science/story/0,,1871842,00.html}}; {{cite web | title = Not a lasting last for the Neandertals | publisher = John Hawks Weblog | date = September 2006 | accessdate = January 1, 2011 | url = http://johnhawks.net/weblog/reviews/neandertals/gorhams_28000_date_2006.html }}
34. ^{{cite journal |vauthors=Soficaru A, Dobos A, Trinkaus E |title=Early modern humans from the Pestera Muierii, Baia de Fier, Romania |journal=Proc. Natl. Acad. Sci. U.S.A. |volume=103 |issue=46 |pages= 17196–17201 |date=November 2006 |pmid=17085588 |pmc=1859909 |doi=10.1073/pnas.0608443103 |bibcode=2006PNAS..10317196S}}
35. ^{{cite journal |vauthors=Green RE, Krause J, Briggs AW, etal |title=A draft sequence of the Neandertal genome |journal=Science |volume=328 |issue=5979 |pages= 710–722 |date=May 2010 |pmid=20448178 |pmc=5100745 |doi=10.1126/science.1188021|bibcode = 2010Sci...328..710G }}
36. ^{{cite journal |vauthors=Duarte C, Maurício J, Pettitt PB, Souto P, Trinkaus E, van der Plicht H, Zilhão J |title=The early Upper Paleolithic human skeleton from the Abrigo do Lagar Velho (Portugal) and modern-human emergence in Iberia |journal=Proceedings of the National Academy of Sciences |volume=96 |issue=13 |pages= 7604–7609 |date=June 1999 |pmid=10377462 |pmc=22133 |doi= 10.1073/pnas.96.13.7604|bibcode = 1999PNAS...96.7604D }}
37. ^{{cite journal |vauthors=Ruzicka J, Hansen EH, Ghose AK, Mottola HA |title=Enzymatic determination of urea in serum based on pH measurement with the flow injection method |journal=Analytical Chemistry |volume=51 |issue=2 |pages= 199–203 |date=February 1979 |pmid=33580 |doi= 10.1021/ac50038a011 |url= }}
38. ^{{Cite journal | doi=10.1038/nature14558| pmid=26098372|pmc = 4537386| title=An early modern human from Romania with a recent Neanderthal ancestor| journal=Nature| volume=524| issue=7564| pages=216–219| year=2015| last1=Fu| first1=Qiaomei| last2=Hajdinjak| first2=Mateja| last3=Moldovan| first3=Oana Teodora| last4=Constantin| first4=Silviu| last5=Mallick| first5=Swapan| last6=Skoglund| first6=Pontus| last7=Patterson| first7=Nick| last8=Rohland| first8=Nadin| last9=Lazaridis| first9=Iosif| last10=Nickel| first10=Birgit| last11=Viola| first11=Bence| last12=Prüfer| first12=Kay| last13=Meyer| first13=Matthias| last14=Kelso| first14=Janet| last15=Reich| first15=David| last16=Pääbo| first16=Svante}}
39. ^{{cite journal |date= October 17, 2012|title= North African Populations Carry the Signature of Admixture with Neandertals|journal= PLoS ONE|doi= 10.1371/journal.pone.0047765|volume=7|issue= 10|pages=e47765|pmid=23082212|pmc=3474783 | last1 = Sánchez-Quinto | first1 = F | last2 = Botigué | first2 = LR | last3 = Civit | first3 = S | last4 = Arenas | first4 = C | last5 = Avila-Arcos | first5 = MC | last6 = Bustamante | first6 = CD | last7 = Comas | first7 = D | last8 = Lalueza-Fox | first8 = C|bibcode= 2012PLoSO...747765S}}
40. ^{{cite journal |date= 23 October 2014|title= Genome sequence of a 45,000-year-old modern human from western Siberia|journal= Nature|volume= 514|issue= 7523|pages= 445–449|doi= 10.1038/nature13810|pmid=25341783|pmc= 4753769|author1= Fu|first1= Q|last2= Li|first2= H|last3= Moorjani|first3= P|last4= Jay|first4= F|last5= Slepchenko|first5= S. M.|last6= Bondarev|first6= A. A.|last7= Johnson|first7= P. L.|last8= Aximu-Petri|first8= A|last9= Prüfer|first9= K|last10= De Filippo|first10= C|last11= Meyer|first11= M|last12= Zwyns|first12= N|last13= Salazar-García|first13= D. C.|last14= Kuzmin|first14= Y. V.|last15= Keates|first15= S. G.|last16= Kosintsev|first16= P. A.|last17= Razhev|first17= D. I.|last18= Richards|first18= M. P.|last19= Peristov|first19= N. V.|last20= Lachmann|first20= M|last21= Douka|first21= K|last22= Higham|first22= T. F.|last23= Slatkin|first23= M|last24= Hublin|first24= J. J.|last25= Reich|first25= D|last26= Kelso|first26= J|last27= Viola|first27= T. B.|last28= Pääbo|first28= S|bibcode= 2014Natur.514..445F|hdl= 10550/42071}}
41. ^Brahic, Catherine. [https://www.newscientist.com/article/dn24988-humanitys-forgotten-return-to-africa-revealed-in-dna.html#.VZy0CvkmNOI "Humanity's forgotten return to Africa revealed in DNA"], The New Scientist (February 3, 2014).
42. ^{{cite web|url=https://www.sciencedaily.com/releases/2016/02/160217140302.htm|title=Neanderthals mated with modern humans much earlier than previously thought, study finds: First genetic evidence of modern human DNA in a Neanderthal individual|website=ScienceDaily|date=17 February 2016|access-date=2016-03-06}}
43. ^"The Combined Landscape of Denisovan and Neanderthal Ancestry in Present-Day Humans", Current Biology, Sankararaman et al., 26, 1241–1247, 2016
44. ^{{cite web|url=http://phys.org/news/2014-02-neanderthals-extinction-modern-humans-emerged.html |title=Neanderthals may have faced extinction long before modern humans emerged |publisher=Phys.org |date=2014-02-24 |accessdate=2017-02-23}}
45. ^{{cite journal | last = Fisher | first = Richard V. |author2=Giovanni Orsi |author3=Michael Ort |author4=Grant Heiken | title = Mobility of a large-volume pyroclastic flow – emplacement of the Campanian ignimbrite, Italy | journal = Journal of Volcanology and Geothermal Research | volume = 56 | issue = 3 | pages =205–220 | date = June 1993 | url = http://volcanology.geol.ucsb.edu/camptuff.htm | doi = 10.1016/0377-0273(93)90017-L | accessdate = 2008-09-20|bibcode = 1993JVGR...56..205F }}
46. ^{{Cite journal|title=Significance of Ecological Factors in the Middle to Upper Paleolithic Transition |journal=Current Anthropology |volume=51 |issue=5 |page=655 |doi=10.1086/656185 |year=2010 |last1=Golovanova |first1=Liubov Vitaliena |last2=Doronichev |first2=Vladimir Borisovich |last3=Cleghorn |first3=Naomi Elansia |last4=Koulkova |first4=Marianna Alekseevna |last5=Sapelko |first5=Tatiana Valentinovna |last6=Shackley |first6=M. Steven }}
47. ^1 2 {{cite news | url = http://news.nationalgeographic.com/news/2010/01/100922-volcanoes-eruptions-neanderthals-science-volcanic-humans/ | title = Volcanoes Killed Off Neanderthals, Study Suggests | last = Than | first = Ker |date = September 22, 2010 | work = National Geographic | accessdate = 23 September 2010}}
48. ^{{cite journal | last1 = Gilligan | first1 = I | year = 2007 | title = Neanderthal extinction and modern human behaviour: the role of climate change and clothing | url = | journal = World Archaeology | volume = 39 | issue = 4| pages = 499–514 | doi=10.1080/00438240701680492}}
49. ^Climate Change Killed Neandertals, Study Says, National Geographic News
50. ^{{cite news | url = https://www.sciencedaily.com/releases/2010/10/101006094057.htm | title = Volcanoes wiped out Neanderthals, new study suggests (ScienceDaily) | date = 7 October 2010 | work = University of Chicago Press Journals | accessdate = 7 October 2010}}
51. ^Neanderthal Apocalypse Documentary film, ZDF Enterprises, 2015. Retrieved 26 January 2016.

External links

  • [https://www.npr.org/templates/story/story.php?storyId=126553081 Hey Good Lookin': Early Humans Dug Neanderthals] – audio report by NPR (6 May 2010)
{{Homo neanderthalensis|state=expanded}}{{DEFAULTSORT:Neanderthal Extinction Hypotheses}}

4 : Neanderthals|Upper Paleolithic|European archaeology|Pleistocene extinctions
随便看

　

开放百科全书收录14589846条英语、德语、日语等多语种百科知识，基本涵盖了大多数领域的百科知识，是一部内容自由、开放的电子版国际百科全书。

　

Copyright © 2023 OENC.NET All Rights Reserved
京ICP备2021023879号 更新时间：2024/9/20 6:10:15