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词条 2009 in archosaur paleontology
释义

  1. Crurotarsans

  2. Non-avian dinosaurs

     Research  Hadrosaur chewing study  New taxa   Aves   Research  Newly described birds  Pterosaurs 

  3. See also

  4. Notes

  5. References

{{Year nav topic5|2009|archosaur paleontology|paleontology|science}}{{Portal|Paleontology|History of science|Dinosaurs}}

The year 2009 in Archosaur paleontology was eventful. Archosaurs include the only living dinosaur group — birds — and the reptile crocodilians, plus all extinct dinosaurs, extinct crocodilian relatives, and pterosaurs. Archosaur paleontology is the scientific study of those animals, especially as they existed before the Holocene Epoch began about 11,700 years ago. The year 2009 in paleontology included various significant developments regarding archosaurs.

This article records new taxa of fossil archosaurs of every kind that have been described during the year 2009, as well as other significant discoveries and events related to paleontology of archosaurs that occurred in the year 2009.

Crurotarsans

  • {{cite journal |last1=Delfino |first1=M. |last2=Smith |first2=T. |year=2009 |title=A reassessment of the morphology and taxonomic status of 'Crocodylus' depressifrons Blainville, 1855 (Crocodylia, Crocodyloidea) based on the Early Eocene remains from Belgium |journal=Zoological Journal of the Linnean Society |volume=156 |issue=1|pages=140–167 |doi=10.1111/j.1096-3642.2008.00478.x}}
Newly named crurotarsans
Name Status Authors Age Unit Location Notes Images
Armadillosuchus[1]

Valid

  • Marinho
  • Carvalho

Late Cretaceous

Adamantina Formation{{flag|Brazil}}

Notosuchian with heavy, armadillo-like body armor consisting of flexible bands and rigid shields

Barcinosuchus[2]

Valid

  • Leardi
  • Pol

Aptian – Albian

Cerro Barcino Formation{{flag|Argentina}}

A peirosaurid

Collilongus[3]

Valid

  • Borsuk−Białynicka
  • Sennikov

Early Olenekian

Czatkowice 1{{flag|Poland}}

Possible rauisuchian

Coringasuchus[4]

Valid

  • Kellner
  • Pinheiro
  • et al.[5]

Early Cenomanian

Alcântara Formation{{flag|Brazil}}
Duerosuchus[6]

Valid

  • Santiago
  • Andrés

Middle Eocene

{{flag|Spain}}
Hypselorhachis[7]

Valid

  • Butler
  • Barrett
  • et al.[8]

Middle Triassic

Manda Beds{{flag|Tanzania}}

Possible ctenosauriscid

Kaprosuchus[9]

Valid

  • Sereno
  • Larson

Upper Cretaceous

Echkar Formation{{flag|Niger}}

Unusual large mahajangasuchid with hypertrophied caniniform teeth and posteriorly projecting horns

Kemkemia[9]

Valid

  • Cau & Maganuco

Cenomanian

Kem Kem Beds{{flag|Morocco}}

Initially identified as a theropod dinosaur,[9] but subsequently discovered to be a crocodyliform.[10]

Khoratosuchus[11]

Valid

  • Lauprasert
  • Cuny
  • et al.[12]

Early Cretaceous

{{flag|Thailand}}

Youngest Mesozoic crocodyliform yet known from Thailand

Laganosuchus [13]

Valid

  • Sereno
  • Larson

Cenomanian

Echkar Formation
Kem Kem Beds{{flag|Niger}}
{{flag|Morocco}}
Miadanasuchus [14]

Valid

  • Simons
  • Buckley

Campanian

Maevarano Formation{{flag|Madagascar}}

A new genus for "Trematochampsa" oblita (Buffetaut & Taquet, 1979)

Morrinhosuchus[15]

Valid

  • Iori
  • Carvalho

Late Cretaceous

Adamantina Formation{{flag|Brazil}}

Notosuchian from Brazil

Penghusuchus[16]

Valid

  • Shan
  • Wu
  • et al.[17]

Late Miocene

{{flag|Taiwan}}

A tomistomine crocodilian.

Polonosuchus[18]

Valid

  • Brusatte
  • Butler
  • et al.[19]

Late Carnian

{{flag|Poland}}

A new genus for "Teratosaurus" silesiacus (Sulej, 2005)

Yacarerani [20]

Valid

  • Novas
  • Pais
  • et al.[21]

Late Cretaceous

Cajones Formation{{flag|Bolivia}}

Non-avian dinosaurs

Research

  • A new study on theropod furculae is published.[22]
  • A "detailed description of the skull and mandible of the Chinese cerapodan ornithischian dinosaur Jeholosaurus shangyuanensis" is published.[23]
  • Knoll, F., Padian, K., and de Ricqles, A. 2009. Ontogenetic change and adult body size of the early ornithischian dinosaur Lesothosaurus diagnosticus: Implications for basal ornithischian taxonomy. Gondwana Research. {{doi|10.1016/j.gr.2009.03.010}}.
  • Matthews, J. C., Brusatte, S. L., Williams, S. A., and Henderson, M. D., 2009, The first Triceratops bonebed and its implications for gregarious behavior Journal of Vertebrate Paleontology v. 29, n. 1, p. 286-290.
  • Williamson, T. E., Carr, T. D., Williams, S. A., and Tremaine, K., 2009, Early ontogeny of pachycephalosaurine squamosals as revealed by juvenile specimens from the Hell Creek Formation, eastern Montana: Journal of Vertebrate Paleontology, v. 29, n. 1, p. 291-294.
  • {{cite journal | last1 = Bittencourt | first1 = J.S. | last2 = Kellner | first2 = A.W.A. | year = 2009 | title = The anatomy and phylogenetic position of the Triassic dinosaur Staurikosaurus pricei Colbert, 1970 | url = | journal = Zootaxa | volume = 2079 | issue = | pages = 1–56 }}
  • {{cite journal | last1 = Chin | first1 = K. | last2 = Hartman | first2 = J.H. | last3 = Roth | first3 = B. | year = 2009 | title = Opportunistic exploitation of dinosaur dung: fossil snails in coprolites from the Upper Cretaceous Two Medicine Formation of Montana | url = | journal = Lethaia | volume = 42 | issue = 2| pages = 185–198 | doi = 10.1111/j.1502-3931.2008.00131.x }}
  • Maidment, S.C.R., and Porro, L.B. 2009. Homology of the palpebral and origin of supraorbital ossifications in ornithischian dinosaurs. Lethaia. {{doi|10.1111/j.1502-3931.2009.00172.x}}.
  • Gates, T.A., and Farke, A.A. 2009. Biostratigraphic and biogeographic implications of a hadrosaurid (Ornithopoda: Dinosauria) from the Upper Cretaceous Almond Formation of Wyoming, USA. Cretaceous Research. {{doi|10.1016/j.cretres.2009.05.001}}.
  • {{cite journal | last1 = Moratalla | first1 = J.J. | last2 = Hernán | first2 = J. | year = 2008 | title = Los Cayos S y D: dos afloramientos con icnitas de saurópodos, terópodos y ornitópodos en el Cretácico Inferior del área de Los Cayos (Cornago, La Rioja, España) | url = | journal = Estudios Geológicos | volume = 64 | issue = 2| pages = 161–173 | doi = 10.3989/egeol.08642.043 }}
  • {{cite journal | last1 = Taylor | first1 = M.P. | last2 = Wedel | first2 = M.J. | last3 = Naish | first3 = D. | year = 2009 | title = Head and neck posture in sauropod dinosaurs inferred from extant animals | url = | journal = Acta Palaeontologica Polonica | volume = 54 | issue = 2| pages = 213–220 | doi=10.4202/app.2009.0007}}

Hadrosaur chewing study

{{Main|2008–2009 hadrosaur chewing study}}

A study titled "Quantitative analysis of dental microwear in hadrosaurid dinosaurs, and the implications for hypotheses of jaw mechanics and feeding" is published by British paleontologists Mark Purnell, Paul Barrett and student Vince Williams. The paper examined the chewing methods and diet of hadrosaurid ("duck billed") dinosaurs from the Late Cretaceous period. The scientists analyzed hundreds of microscopic scratches on the teeth of a fossilized Edmontosaurus jaw, and believe they determined exactly how a hadrosaur broke down and ate its food, which had previously eluded researchers.

The study found hadrosaurs had a unique way of eating unlike any creature living today. In contrast to a flexible lower jaw joint prevalent in today's mammals, hadrosaurs had a unique hinge between the upper jaws and the rest of its skull. The team found the dinosaur's upper jaws pushed outwards and sideways while chewing, as the lower jaw slid against the upper teeth.

The study also concluded that hadrosaurs likely grazed on horsetails and vegetation close to the ground, rather than browsing higher-growing leaves and twigs. However, Purnell said these conclusions were less secure than the more conclusive evidence regarding the motion of teeth while chewing. Previous studies found contradictory conclusions, and the issue remains a subject of debate.

The findings were published on June 30, 2009 in the journal, The Proceedings of the National Academy of Sciences. Purnell said no previous study had ever employed this method of analyzing microscopic teeth scratches, and that the method could be used to study other areas of scientific research.

New taxa

Data courtesy of George Olshevky's dinosaur genera list.[24] ~44 dinosaur genera were erected in 2009.

Albalophosaurus[28]Angulomastacator[32]

Valid

Arenysaurus[33]

Valid

Australovenator[35]

Valid

Barrosasaurus[38]Elrhazosaurus[41]

Valid

Jintasaurus[45]

Valid

Leshansaurus[48]

Valid

Owenodon[41]

Valid

Qiaowanlong[45]Raptorex[58]Sinotyrannus[66]

Valid

Spinophorosaurus[69]Tatankacephalus[71]

Valid

Tawa[72]

Valid

Tethyshadros[74]

Valid

Tianyulong[75]

Valid

Wintonotitan[40]

Valid

"Xinghesaurus"

Nomen nudumZanabazar[79]

Valid

Name Status Authors Discovery year Age Unit LocationNotes Images
Adeopapposaurus[25] Valid
  • Ricardo N. Martínez
Cañón del Colorado Formation
  • {{Flag|Argentina}}
Aerosteon[26] Valid
  • Sereno
  • R. N. Martinez
  • et al.[27]
Rio Colorado Formation
  • {{Flag|Argentina}}
Valid
  • Ohashi
  • Barrett
Kuwajima Formation
  • {{Flag|Japan}}
Albertonykus[29] Valid
  • Longrich
  • Currie

lower Maastrichtian

Horseshoe Canyon Formation
  • {{Flag|Canada}} ({{Flag|Alberta}})
Anchiornis[30] Valid
  • Xu X.
  • Zhao Q.
  • et al.[31]
Tiaojishan Formation
  • {{Flag|China}}
  • J. R. Wagner
  • Lehman
Aguja Formation
  • {{Flag|USA}} ({{Flag|Texas}})
  • Xabier Pereda-Suberbiolaa
  • José Ignacio Canudob
  • et al.[34]
  • {{Flag|Spain}}
A Spanish Lambeosaurine.
  • Hocknull
  • White
  • et al.[38]
Winton Formation
  • {{Flag|Australia}}
An Australian Megaraptor. Specimen named "Banjo"
Baotianmansaurus[36]

Valid

  • Zhang, X.
  • Lü, J.
  • et al.[37]
Gaogou Formation
  • {{Flag|China}}
Valid
  • Salgado
  • Coria
Anacleto Formation
  • {{Flag|Argentina}}
Ceratonykus[39]

Valid

  • Alifanov
  • Barsbold
Barun Goyot Formation
  • {{Flag|Mongolia}}
Diamantinasaurus[40]

Valid

  • Hocknull
  • White
  • et al.[38]
Winton Formation
  • {{Flag|Australia}}
An Australian Titanosaur.
  • Galton
Elrhaz Formation{{Flag|Niger}}
Helioceratops[42]

Valid

  • Jin
  • Chen
  • et al.[43]
Quantou Formation
  • {{Flag|China}}
Hesperonychus[44]

Valid

  • Longrich
  • Currie
Dinosaur Park Formation
  • {{Flag|Canada}}

Smallest known dinosaur from North America.

  • You
  • Li
Xinminpu Group
  • {{Flag|China}}
Kinnareemimus[46]

Valid

  • Buffetaut
  • Suteethorn
  • Tong
Sao Khua Formation
  • {{Flag|Thailand}}
Kol[47]

Valid

  • Turner
  • Nesbitt
  • Norell
Djadochta Formation
  • {{Flag|Mongolia}}
  • Li
  • Peng
  • et al.[49]
Shangshaximiao Formation
  • {{Flag|China}}
Levnesovia[50] Valid
  • Sues
  • Averianov
Bissekty Formation
  • {{Flag|Uzbekistan}}

The oldest Hadrosauroidean

Limusaurus[51] Valid
  • Xu
Shishugou Formation
  • {{Flag|China}}

The first Asian ceratosaur to be discovered

Luoyanggia[52] Valid
  • Xu
  • et al.[53]
Mangchuan Formation
  • {{Flag|China}}

An oviraptorosaur

Malarguesaurus[54] Valid
  • González Riga
  • Previtera
  • Pirrone
Portezuelo Formation
  • {{Flag|Argentina}}
Minotaurasaurus[55] Valid
  • Clifford A. Miles
  • Clark J. Mikes
  • {{Flag|Mongolia}}
Synonym of Tarchia.
Miragaia[56] Valid
  • Mateus
  • Maidment
  • Christiansen
Sobral Unit
  • {{Flag|Portugal}}

Long-necked stegosaur.

  • Galton
Purbeck Limestone
  • {{Flag|UK}}
Panphagia[57] Valid
  • Martinez
  • Alcober
Ischigualasto Formation
  • {{Flag|Argentina}}

One of the most basal known sauropodomorph.[57]

Valid
  • You
  • Li
Xinminpu Group
  • {{Flag|China}}
  • Sereno
  • Brusatte
  • et al.[59]
Yixian Formation
  • {{Flag|China}}
Tyrannosauroidea
Ruyangosaurus[60]

Valid

  • Lu
  • Xu
  • et al.[61]
Mangchuan Formation
  • {{Flag|China}}
Shaochilong[62]

Valid

  • Brusatte
  • Benson
  • et al.[63]
Ulansuhai Formation
  • {{Flag|China}}
Shidaisaurus[64]

Valid

  • Wu
  • Currie
  • et al.[65]
Upper Lufeng Formation
  • {{Flag|China}}
  • Ji
  • Ji
  • Zhang
Jiufotang Formation
  • {{Flag|China}}
Skorpiovenator[67] Valid
  • Canale
  • Scanferla
  • et al.[68]
Huincul Formation
  • {{Flag|Argentina}}
  • Remes
  • Ortega
  • et al.[70]
Irhazer Group
  • {{Flag|Niger}}
Sauropoda
  • Parsons
  • Parsons
Cloverly Formation
  • {{Flag|USA}} ({{Flag|Montana}})
  • Nesbitt
  • Smith
  • et al.[73]
Chinle Formation
  • {{Flag|USA}} ({{Flag|New Mexico}})
  • Dalla Vecchia
Liburnia Formation
  • {{Flag|Italy}}
  • Zheng
  • You
  • et al.[76]
Tiaojishan Formation
  • {{Flag|China}}
  • Hocknull
  • White
  • et al.[77]
Winton Formation
  • {{Flag|Australia}}
Xianshanosaurus[52] Valid
  • Xu
  • et al.[53]
Mangchuan Formation
  • {{Flag|China}}

A sauropod

  • Hasegawa
  • Carpenter
  • et al.[78]
Name published without scientific description in Japanese guidebook "Dinosaur Expo 2009: The Miracle of Deserts"
  • Norell
  • Makovicky
  • et al.[80]
Nemegt Formation
  1. {{Flag|Mongolia}}

Aves

Research

  • Anfinson, O.A., Lockley, M.G., Kim, S.H., Kim, K.S., and Kim, J.Y. 2009. First report of the small bird track Koreanaornis from the Cretaceous of North America: implications for avian ichnotaxonomy and paleoecology. Cretaceous Research. {{doi|10.1016/j.cretres.2009.02.001}}.
  • Lockley, M., Chin, K., Houck, K., Matsukawa, M., and Kukihara, R. 2009. New interpretations of Ignotornis, the first-reported Mesozoic avian footprints: implications for the paleoecology and behavior of an enigmatic Cretaceous bird. Cretaceous Research. {{doi|10.1016/j.cretres.2009.04.001}}.
  • {{cite journal | last1 = Bell | first1 = A. | last2 = Everhart | first2 = M.J. | year = 2009 | title = A new specimen of Parahesperornis (Aves: Hesperornithiformes) from the Smoky Hill Chalk (Early Campanian) of western Kansas | url = | journal = Transactions of the Kansas Academy of Science | volume = 112 | issue = 1/2| pages = 7–14 | doi=10.1660/062.112.0202}}

Newly described birds

Alamitornis minutus[81]

Gen. nov. et Sp. nov.

Valid

Federico L. AgnolinAgustín G. MartinelliLate CretaceousLos Alamitos Formation
  • {{Flag|Argentina}}

Placed in ?Patagopterygiformes by Agnolin and Martinelli.

Australotadorna alecwilsoni[82]

Gen. nov. et Sp. nov.

Valid

Trevor H. Worthy

Late Oligocene

Etadunna Formation
  • {{Flag|Australia}}

An Anatidae.

Bonasa nini[83]

Sp. nov.

Valid

Antonio Sánchez Marco

Early Pleistocene

Lower Elefante, TE 13 layer

  • {{Flag|Spain}}

A Phasianidae.

Calidris janossyi[84]

Sp. nov.

Valid

Jenö Kessler

Late Miocene

MN 13
  • {{Flag|Hungary}}

Originally described as a member of Scolopacidae belonging to the genus Calidris;[84] Zelenkov, Volkova and Gorobets (2016) reinterpreted it as a member of Turnicidae and transferred it to the genus Ortyxelos.[85]

Calonectris kurodai[86]

Sp. nov.

Valid

Storrs L. Olson

Middle Miocene

Calvert Formation
  • {{Flag|USA}} ({{Flag|Virginia}})

A Procellariidae, the smallest of the genus.

Cariama santacrucensis[87]

Sp. nov.

Disputed

Jorge I. NoriegaSergio F. VizcainoSusana Bargo

Early-Middle Miocene

Estancia La Costa Member,

Santa Cruz Formation
  • {{Flag|Argentina}}

Federico L. Agnolin, 2009, made it the type species of the separate genus Noriegavis Agnolin, 2009.[88] Originally interpreted as a seriema; Noriega & Mayr (2017) reinterpreted it as a member of the falconid genus Thegornis of uncertain specific assignment, on the basis of a reexamination of the holotype specimen.[89]

Charadrius lambrechti.[84]

Sp. nov.

Valid

Jenö Kessler

Late Miocene

MN 13
  • {{Flag|Hungary}}

A Scolopacidae.

Clangula matraensis[90]

Sp. nov.

Valid

Jenö Kessler

Middle Miocene

MN 6-8

  • {{Flag|Hungary}}

An Anatidae.

Confuciusornis feducciai[91]

Sp. nov.

Disputed

Zuhui ZhangChunling GaoQingjin MengJinyuan LiuLianhai HouGuangmei ZhengEarly CretaceousYixian Formation
  • {{Flag|China}}

A member of the family Confuciusornithidae. Considered to be a junior synonym of Confuciusornis sanctus by Wang, O'Connor & Zhou (2018).[92]

Cygnopterus neogradiensis[93]

Sp. nov.

Valid

Jenö KesslerJános Hír

Middle Miocene

Sajóvölgy Formation,

MN 7-8

  • {{Flag|Hungary}}

An Anatidae.

Diomedeoides harmati[94]

Sp. nov.

Valid

Jenö Kessler

Early Oligocene

MP 24
  • {{Flag|Hungary}}

A Diomedeoididae Fischer, 1985, placed in Rupelornis van Beneden, 1871 by Gerald Mayr and Thierry Smith, 2012

Egretta polgardiensis[90]

Sp. nov.

Disputed

Jenö Kessler

Late Miocene

MN 13
  • {{Flag|Hungary}}

Originally described as a member of Ardeidae and a species of Egretta; however, Zelenkov (2017) considered this species to be a junior synonym of the barn-owl species Tyto campiterrae Jánossy (1991).[108]

Elbretornis bonapartei[95]

Gen. nov. et Sp. nov.

Valid

Cyril A. WalkerGareth J. DykeMaastrichtianLecho Formation
  • {{Flag|Argentina}}

An Enantiornithes Walker, 1981, Euenantiornithes Chiappe, 2002, this is the type species of the new genus.

Eoanseranas handae[96]

Gen. nov. et Sp. nov.

Valid

Trevor H. WorthyJohn D. Scanlon

Late Oligocene – Early Miocene

Riverleigh World Heritage Property

  • {{Flag|Australia}}

An Anseranatidae, this is the type species of the new genus.

Gansugyps linxiaensis[97]

Gen. nov. et Sp. nov.

Valid

Zhang ZihuiZheng XiaotingZheng GuangmeiHou Lianhai

Late Miocene

Upper Liushu Formation

  • {{Flag|China}}

An Accipitridae, this is the type species of the new genus.

Heliadornis minor[90]

Sp. nov.

Valid

Jenö Kessler

Late Pliocene

MN 15-16

  • {{Flag|Slovakia}}

A Phaethontidae.

Heliornis sumeghensis[84]

Sp. nov.

Valid

Jenö Kessler

Late Miocene

MN 11-12

  • {{Flag|Hungary}}

A Heliornithidae

Jianchangornis microdonta[98]

Gen. nov. et Sp. nov.

Valid

Zhou ZhongheZhang FuchengLi ZhihengEarly CretaceousAlbianJiufotang Formation
  • {{Flag|China}}

A basal Ornithurae Haeckel, 1866, this is the type species of the new genus.

Martinavis minor[95]

Sp. nov.

Valid

Cyril A. WalkerGareth J. DykeLate CretaceousMaastrichtianLecho Formation
  • {{Flag|Argentina}}

An Enantiornithes Walker, 1981, Euenantiornithes Chiappe, 2002.

Martinavis saltariensis[95]

Sp. nov.

Valid

Cyril A. WalkerGareth J. DykeLate CretaceousMaastrichtianLecho Formation
  • {{Flag|Argentina}}

An Enantiornithes Walker, 1981, Euenantiornithes Chiappe, 2002.

Martinavis whetstonei[95]

Sp. nov.

Valid

Cyril A. WalkerGareth J. DykeLate CretaceousMaastrichtianLecho Formation
  • {{Flag|Argentina}}

An Enantiornithes Walker, 1981, Euenantiornithes Chiappe, 2002.

Megalocoturnix cordoni[83]

Gen. nov. et Sp. nov.

Valid

Antonio Sánchez Marco

Early Pliocene

Ruscinian,

MN 15
  • {{Flag|Spain}}

A Phasianidae, this is the type species of the new genus.

Megapaloelodus peiranoi[99]

Sp. nov.

Valid

Federico L. Agnolin

Late Miocene

Andalhualá Formation
  • {{Flag|Argentina}}

A Phoenicopteriformes, Palaelodidae Stejneger, 1885.

Mergus minor[90]

Sp. nov.

Valid

Jenö Kessler

Middle Miocene

MN 6-8

  • {{Flag|Hungary}}

An Anatidae.

Miocepphus blowi[100]

Gen. nov. et Sp. nov.

Valid

Erik WijnkerStorrs L. Olson

Late Miocene

Eastover Formation;

Breda Formation Mill

  • {{Flag|USA}} ({{Flag|Virginia}})
  • {{Flag|Netherlands}}

An Alcidae.

Miocepphus bohaskai[100]

Sp. nov.

Valid

Erik WijnkerStorrs L. Olson

Late Miocene

Eastover Formation;

Breda Formation Mill

  • {{Flag|USA}} ({{Flag|Maryland}} and {{Flag|Virginia}})
  • {{Flag|Netherlands}}

An Alcidae.

Miocepphus mergulellus[100]

Sp. nov.

Valid

Erik WijnkerStorrs L. Olson

Late Miocene

Eastover Formation
  • {{Flag|USA}} ({{Flag|North Carolina}})

An Alcidae.

Palaeocryptonyx novaki[83]

Sp. nov.

Valid

Antonio Sánchez Marco

Late Pliocene

Villanyan,

MN 16
  • {{Flag|Spain}}

A Phasianidae.

Paracoracias occidentalis[101]

Gen. nov. et Sp. nov.

Valid

Julia A. ClarkeDaniel T. KsepkaN. Adam SmithMark A. NorellEoceneGreen River Formation
  • {{Flag|USA}} ({{Flag|Wyoming}})

A stem Coraiiformes.

Pinpanetta fromensis[82]

Sp. nov.

Valid

Trevor H. Worthy

Late Oligocene

Etadunna Formation
  • {{Flag|Australia}}

An Anatidae.

Pinpanetta tedfordi[82]

Gen. nov. et Sp. nov.

Valid

Trevor H. Worthy

Late Oligocene

Etadunna Formation
  • {{Flag|Australia}}

An Anatidae, this is the type species of the new genus.

Pinpanetta vickersrichae[82]

Sp. nov.

Valid

Trevor H. Worthy

Late Oligocene

Etadunna Formation
  • {{Flag|Australia}}

An Anatidae.

Podiceps csarnotatus[90]

Sp. nov.

Valid

Jenö Kessler

Late Pliocene

MN 15-16

  • {{Flag|Hungary}}

A Podicipedidae.

Porzana kretzoii[84]

Sp. nov.

Valid

Jenö Kessler

Late Miocene

MN 13
  • {{Flag|Hungary}}

A member of the family Rallidae. Originally described as a species Porzana; Zelenkov (2017) transferred this species to the genus Zapornia.[102]

Porzana matraensis[84]

Sp. nov.

Valid

Jenö Kessler

Middle Miocene

MN 6-8

  • {{Flag|Hungary}}

A Rallidae.

Primozygodactylus eunjooae[103]

Sp. nov.

Valid

Gerald MayrNikita V. Zelenkov

Middle Eocene

MP 11
  • {{Flag|Germany}}

A Zygodactylidae Brodkorb, 1971.

Pseudocepphus teres[100]

Gen. nov. et Sp. nov.

Valid

Erik WijnkerStorrs L. Olson

Late Miocene

St. Marys Formation
  • {{Flag|USA}} ({{Flag|Maryland}})

An Alcidae, this is the type species of the new genus.

Rapaxavis pani[104]

Gen. nov. et Sp. nov.

Valid

Eric M. MorschhauserDavid J. VarricchioGao ChunlingLiu JinyuanWang XuriCheng XiadongMeng QingjinEarly CretaceousJiufotang Formation
  • {{Flag|China}}

An Enantiornithes Walker, 1981, Longipterygidae Zhang, Zhou, Hou et Gu, 2000, this is the type species of the new genus.

Rupephaps taketake[105]

Gen. nov. et Sp. nov.

Valid

Trevor H. WorthySuzanne J. HandJennifer P. WorthyAlan J. D. TennysonR. Paul Scofield

Early Miocene,

AltonianBannockburn Formation
  • {{Flag|New Zealand}}

A Columbidae, this is the type species of the new genus.

Sapeornis angustis[106]

Sp. nov.

Valid

Pauline ProviniZhou ZhongheZhang Fucheng

Early Cretaceous,

AptianJiufotang Formation
  • {{Flag|China}}

A Sapeornithidae Zhou et Zhang, 2006.

Selenornis steendorpensis[107]

Sp. nov.

Valid

Gerald Mayr

Early Oligocene,

MP ?23-24

Boom Formation
  • {{Flag|Belgium}}

A Tytonidae, Selenornithinae Mourer-Chauviré, 1987.

Shanweiniao cooperorum[108]

Gen. nov. et Sp. nov.

Valid

Jingmai K. O'ConnorXuri WangLuis M. ChiappeChunling GaoQingjin MengXiaodong ChengJinyuan LiuEarly CretaceousYixian Formation
  • {{Flag|China}}

An Enantiornithes Walker, 1981, Familia Longipterygidae Zhang, Zhou, Hou et Gu, 2000, this is the type species of the new genus.

Talpanas lippa[109]

Gen. nov. et Sp. nov.

Valid

Storrs L. OlsonHelen F. JamesHoloceneKauai
  • {{Flag|USA}} ({{Flag|Hawaii}})

An Anatidae, Kaua'i Mole Duck, a Hawaiian duck that probably lived a kiwi-like lifestyle, this is the type species of the new genus.

Tologuica aurorae[110]

Gen. nov. et Sp. nov.

Valid

Nikita V. ZelenkovEvgeny N. Kurochkin

Middle Miocene

Ööshin Formation
  • {{Flag|Mongolia}}

A Phasianidae, this is the type species of the new genus.

Tologuica karhui[110]

Sp. nov.

Valid

Nikita V. ZelenkovEvgeny N. Kurochkin

Middle Miocene

Ööshin Formation
  • {{Flag|Mongolia}}

A Phasianidae.

Name Novelty Status Authors Age Unit Location Notes Images

Pterosaurs

  • {{cite journal | last1 = Lü | first1 = J. | year = 2009 | title = A new non-pterodactyloid pterosaur from Qinglong County, Hebei Province of China | url = | journal = Acta Geologica Sinica (English Edition) | volume = 83 | issue = 2| pages = 189–199 | doi = 10.1111/j.1755-6724.2009.00062.x }}
  • Vullo, R., and Neraudeau, D., 2009, Pterosaur remains from the Cenomanian (Late Cretaceous) Paralic Deposits of Charentes, Western France Journal of Vertebrate Paleontology, v. 29, n. 1, p. 277-282.
Carniadactylus[111]

Valid

  • Dalla Vecchia

Late Triassic

  • {{Flag|Italy}}

A campylognathoidid with a wingspan of about 70 cm.

Changchengopterus[112]

Valid

Middle Jurassic

Tiaojishan Formation
  • {{flag|China}}

A primitive long-tailed pterosaur related to Dorygnathus.

Ningchengopterus[113]

Valid

Early Cretaceous

Yixian Formation
  • {{flag|China}}

A pterodactyloid known from a juvenile specimen whose fossils preserved details of the flight membrane and fur.

Wukongopterus[114]

Valid

  • Wang
  • Kellner
  • Jiang
  • Meng

Late Jurassic

Daohugou Beds
  • {{flag|China}}

A primitive long tailed pterosaur. The type specimen shows evidence of the animal having broken its shin while alive.

Newly named pterosaurs
Name Status Authors Age Unit LocationNotesImages

See also

  • 2010 in archosaur paleontology
  • 2011 in archosaur paleontology
  • 2012 in archosaur paleontology
  • 2013 in archosaur paleontology

Notes

1. ^{{Cite journal |author1=Marinho, T.S. |author2=Carvalho, I.S. |year= 2009 |title= An armadillo-like sphagesaurid crocodyliform from the Late Cretaceous of Brazil |journal= Journal of South American Earth Sciences |volume=27 |issue=1 |pages=36–41 |doi= 10.1016/j.jsames.2008.11.005|bibcode = 2009JSAES..27...36M }}
2. ^{{Cite journal|author1=Juan Martín Leardi |author2=Diego Pol |year=2009 |title=The first crocodyliform from the Chubut Group (Chubut Province, Argentina) and its phylogenetic position within basal Mesoeucrocodylia |url= |journal=Cretaceous Research |volume=30 |issue=6 |pages=1376–1386 |doi=10.1016/j.cretres.2009.08.002}}
3. ^{{Cite journal|author1=Magdalena Borsuk−Białynicka |author2=Andriej G. Sennikov |year=2009|title=Archosauriform postcranial remains from the Early Triassic karst deposits of southern Poland|journal=Paleontologica Polonica|volume=65|pages=283–328|url=http://palaeontologia.pan.pl/PP65/PP65_283-328.pdf| accessdate= 13 August 2010 }}
4. ^{{Cite journal | author= Kellner, A.W.A. |author2=Pinheiro, A.E.P. |author3=Azevedo, S.A.K. |author4=Henriques, D.D.R. |author5=de Carvalho, L.B. |author6=Oliveira, G.R. |year= 2009 |title= A new crocodyliform from the Alcântara Formation (Cenomanian), Cajual Island, Brazil |journal= Zootaxa |volume= 2030 |pages=49–58}}
5. ^Kellner, Pinheiro, Azevedo, Henriques, de Carvalho, Oliveira.
6. ^{{cite journal |last=Santiago |first=L.A. |author2=and Andrés, L.A. |year=2009 |title=Nuevo género de Crocodylia del Eoceno medio de la Península Ibérica (Zamora, España): Duerosuchus piscator nov. gen., nov. sp |journal=Studia Geologica Salmanticensia |volume=45 |issue=2 |pages=149–173}}
7. ^{{Cite journal |author1=Butler, R.J. |author2=Barrett, P.M. |author3=Abel, R.L. |author4=Gower, D.J. |year=2009 |title=A Possible Ctenosauriscid Archosaur from the Middle Triassic Manda Beds of Tanzania |journal=Journal of Vertebrate Paleontology |volume=29 |issue=4 |pages=1022–1031 |doi=10.1671/039.029.0404}}
8. ^Butler, Barrett, Abel, Gower.
9. ^{{cite journal |first=Andrea |last=Cau |first2=Simone |last2=Maganuco |title=A new theropod dinosaur, represented by a single unusual caudal vertebra from the Kem Kem Beds (Cretaceous) of Morocco |journal=Atti Soc. It. Sci. Nat. Museo Civ. Stor. Nat. Milano |volume=150 |issue=II |pages=239–257 |year=2009}}
10. ^Lio, G., Agnolin, F., Cau, A. and Maganuco, S. (2012). "Crocodyliform affinities for Kemkemia auditorei Cau and Maganuco, 2009, from the Late Cretaceous of Morocco." Atti della Società Italiana di Scienze Naturali e del Museo di Storia Naturale di Milano, 153 (I), s. 119–126.
11. ^Lauprasert, K.; Cuny, G.; Thirakhupt, K. and Suteethorn, V. 2009. Khoratosuchus jintasakuli gen. et sp. nov., an advanced neosuchian crocodyliform from the Early Cretaceous (Aptian-Albian) of NE Thailand. In E. Buffetaut, G. Cuny, J. Le Loeuff, V. Suteethorn (eds.), Late Palaeozoic and Mesozoic Ecosystems in SE Asia. The Geological Society of London, Special Publication 315:175-187.
12. ^Lauprasert, Cuny, Thirakhupt, Suteethorn.
13. ^{{cite journal |last=Sereno |first=P. C. |author2=Larsson, H. C. E. |year=2009 |title=Cretaceous crocodyliforms from the Sahara |journal=ZooKeys |issue=2009 |pages=1–143 |doi=10.3897/zookeys.28.325}}
14. ^{{cite journal |author1=Erin L. Rasmusson Simons |author2=Gregory A. Buckley |year=2009 |title=New Material of "Trematochampsa" Oblita (Crocodyliformes, Trematochampsidae) from the Late Cretaceous of Madagascar |journal=Journal of Vertebrate Paleontology |volume=29 |issue=2 |pages=599–604 |doi=10.1671/039.029.0224}}
15. ^{{cite journal |last=Iori |first=F.V. |author2=Carvalho, I.S. |year=2009 |title=Morrinhosuchus luziae, um novo Crocodylomorpha Notosuchia da Bacia Bauru, Brasil |journal=Revista Brasileira de Geociências |volume=39 |issue=4 |pages=717–725 |url=http://ojs.c3sl.ufpr.br/ojs2/index.php/rbg/article/viewFile/12839/11539| accessdate= 7 August 2010 |doi=10.25249/0375-7536.2009394717725 }}
16. ^{{cite journal |author=Shan, Hsi-yin |author2=Wu, Xiao-chun |author3=Cheng, Yen-nien |author4=Sato, Tamaki |year=2009 |title=A new tomistomine (Crocodylia) from the Miocene of Taiwan |journal=Canadian Journal of Earth Sciences |volume=46 |issue=7 |pages=529–555 |doi=10.1139/E09-036 |url=http://rparticle.web-p.cisti.nrc.ca/rparticle/AbstractTemplateServlet?calyLang=eng&journal=cjes&volume=46&year=2009&issue=7&msno=e09-036|bibcode = 2009CaJES..46..529S | accessdate= 15 September 2010 }}
17. ^Shan, Wu, Cheng, Sato.
18. ^{{cite journal |last1=Brusatte |first1=Stephen L. |last2=Butler |first2=Richard J. |last3=Sulej |first3=Tomasz |last4=Niedźwiedzki |first4=Grzegorz |author5=Brusatte, Stephen L. |author6=Butler, Richard J. |author7=Sulej, Tomasz |author8=Niedźwiedzki, Grzegorz |year=2009 |title=The taxonomy and anatomy of rauisuchian archosaurs from the Late Triassic of Germany and Poland |journal=Acta Palaeontologica Polonica |volume=54 |issue=2 |pages=221–230 |doi=10.4202/app.2008.0065 |url=http://app.pan.pl/archive/published/app54/app54-221.pdf| accessdate= 13 August 2010 }}
19. ^Brusatte, Butler, Sulej, Niedźwiedzki.
20. ^{{cite journal |last=Novas |first=F. E. |author2=Pais, D. F. |author3=Pol, D. |author4=Carvalho, I. D. S. |author5=Mones, A. |author6=Scanferla, A. |author7=Riglos, M. S. |year=2009 |title=Bizarre notosuchian crocodyliform with associated eggs from the Upper Cretaceous of Bolivia |journal=Journal of Vertebrate Paleontology |volume=29 |issue=4 |pages=1316–1320 |doi=10.1671/039.029.0409}}
21. ^Novas, Pais, Pol, Carvalho, Mones, Scanferla, Riglos.
22. ^Nesbitt, S.J., Turner, A.H., Spaulding, M., Conrad, J.L., and Norell, M.A. 2009. The theropod furcula. Journal of Morphology. {{doi|10.1002/jmor.10724}}.
23. ^{{cite journal | last1 = Barrett | first1 = P.M. | last2 = Han | first2 = F.-L. | year = 2009 | title = Cranial anatomy of Jeholosaurus shangyuanensis (Dinosauria: Ornithischia) from the Early Cretaceous of China | url = | journal = Zootaxa | volume = 2072 | issue = | pages = 31–55 }}
24. ^{{cite web| last = Olshevsky| first = George | title = Dinogeorge's Dinosaur Genera List | url = http://www.polychora.com/dinolist.html| accessdate = 2009-01-05 }}
25. ^{{cite journal |last=Martínez |first=Ricardo N. |year=2009 |title=Adeopapposaurus mognai, gen. et sp. nov (Dinosauria: Sauropodomorpha), with comments on adaptations of basal sauropodomorpha |journal=Journal of Vertebrate Paleontology |volume=29 |issue=1 |pages=142–164 |doi=10.1671/039.029.0102}}
26. ^{{cite journal |vauthors=Sereno PC, Martinez RN, Wilson JA, Varricchio DJ, Alcober OA |date=Sep 2008|title=Evidence for Avian Intrathoracic Air Sacs in a New Predatory Dinosaur from Argentina|journal=PLoS ONE|volume=3|issue=9|page=e3303|pmid=18825273 |pmc=2553519|doi=10.1371/journal.pone.0003303|bibcode = 2008PLoSO...3.3303S |editor1-last=Kemp |editor1-first=Tom |display-authors=etal}}
27. ^Sereno, R. N. Martinez, J. A. Wilson, Varricchio, Alcober.
28. ^{{cite journal | last1 = Ohashi | first1 = T. | last2 = Barrett | first2 = P. M. | year = 2009 | title = A new ornithischian dinosaur from the Lower Cretaceous Kuwajima Formation of Japan | url = | journal = Journal of Vertebrate Paleontology | volume = 29 | issue = 3| pages = 748–757 | doi=10.1671/039.029.0306}}
29. ^{{cite journal |last=Longrich |first=Nicholas R. |author2=Currie, Philip J. |year=2009 |title=Albertonykus borealis, a new alvarezsaur (Dinosauria: Theropoda) from the Early Maastrichtian of Alberta, Canada: Implications for the systematics and ecology of the Alvarezsauridae |journal=Cretaceous Research |volume=30 |issue=1 |pages= 239–252|doi=10.1016/j.cretres.2008.07.005 }}
30. ^Xu, X., Zhao, Q., Norell, M., Sullivan, C., Hone, D., Erickson, G., Wang, X., Han, F. and Guo, Y. (2009). "A new feathered maniraptoran dinosaur fossil that fills a morphological gap in avian origin." Chinese Science Bulletin, six pages, accepted November 15, 2008.
31. ^Xu X., Zhao Q., Norell, C. Sullivan, Hone, Erickson, Wang X. L., Han F., Guo.
32. ^{{cite journal | last1 = Wagner | first1 = J.R. | last2 = Lehman | first2 = T.M. | year = 2009 | title = An Enigmatic New Lambeosaurine Hadrosaur (Reptilia: Dinosauria) from the Upper Shale Member of the Campanian Aguja Formation of Trans-Pecos Texas | url = | journal = Journal of Vertebrate Paleontology | volume = 29 | issue = 2| pages = 605–611 | doi=10.1671/039.029.0208}}
33. ^{{cite journal |last=Pereda-Suberbiola |first=Xabier |author2=José Ignacio Canudo |author3=Penélope Cruzado-Caballero |author4=José Luis Barco |author5=Nieves López-Martínez |author6=Oriol Oms |author7=José Ignacio Ruiz-Omeñaca |year=2009 |title=The last hadrosaurid dinosaurs of Europe: A new lambeosaurine from the Uppermost Cretaceous of Aren (Huesca, Spain) |journal=Comptes Rendus Palevol |volume=8 |issue=6 |pages=559–572 |doi=10.1016/j.crpv.2009.05.002}}
34. ^Xabier Pereda-Suberbiolaa, José Ignacio Canudob, Penélope Cruzado-Caballerob, José Luis Barcoc, Nieves López-Martínezd, Oriol Omse, José Ignacio Ruiz-Omeñaca.
35. ^{{cite journal |last=Molnar |first=Ralph E. |author2=Flannery, Timothy F. |author3=Rich, Thomas H.V. |year=1981 |title=An allosaurid theropod dinosaur from the Early Cretaceous of Victoria, Australia |journal=Alcheringa |volume=5 |pages=141–146 |doi=10.1080/03115518108565427 |issue=2 }}
36. ^{{cite journal | last1 = Zhang | first1 = X. | last2 = Lü | first2 = J. | last3 = Xu | first3 = L. | last4 = Li | first4 = J. | last5 = Yang | first5 = L.K. | last6 = Hu | first6 = W. | last7 = Jia | first7 = S. | last8 = Ji | first8 = Q. | last9 = Zhang | first9 = C. | year = 2009 | title = A new sauropod dinosaur from the Late Cretaceous Gaogou Formation of Nanyang, Henan Province | url = | journal = Acta Geologica Sinica (English Edition) | volume = 83 | issue = 2| pages = 212–221 | doi = 10.1111/j.1755-6724.2009.00032.x }}
37. ^Zhang, X., Lü, J., Xu, L., Li, J., Yang, L.K., Hu, W., Jia, S., Ji, Q. Zhang, C.
38. ^{{cite journal | last1 = Salgado | first1 = L. | last2 = Coria | first2 = R. A. | year = 2009 | title = Barrosasaurus casamiquelai gen. et sp. nov., a new titanosaur (Dinosauria, Sauropoda) from the Anacleto Formation (Late Cretaceous: early Campanian) of Sierra Barrosa (Neuquén, Argentina) | url = | journal = Zootaxa | volume = 2222 | issue = | pages = 1–16 }}
39. ^{{cite journal | last1 = Alifanov | first1 = V.R. | last2 = Barsbold | first2 = R. | year = 2009 | title = Ceratonykus oculatus gen. et sp. nov., a new dinosaur (?Theropoda, Alvarezsauria) from the Late Cretaceous of Mongolia" (in English) | url = | journal = Paleontological Journal | volume = 43 | issue = 1| pages = 86–99 | doi = 10.1134/S0031030109010109 }}
40. ^{{cite journal |last=Hocknull |first=Scott A. |author2=White, Matt A. |author3= Tischler, Travis R. |author4=Cook, Alex G. |author5=Calleja, Naomi D. |author6=Sloan, Trish |author7=Elliott, David A. |year=2009 |title=New Mid-Cretaceous (Latest Albian) Dinosaurs from Winton, Queensland, Australia |journal=PLoS ONE |volume=4 |issue=7 |doi=10.1371/journal.pone.0006190 |pages=e6190 |pmid=19584929 |pmc=2703565 |bibcode = 2009PLoSO...4.6190H |editor1-last=Sereno |editor1-first=Paul }}
41. ^{{cite journal |last=Galton |first=Peter M. |authorlink=Peter Galton |year=2009 |title=Notes on Neocomian (Late Cretaceous) ornithopod dinosaurs from England – Hypsilophodon, Valdosaurus, "Camptosaurus", "Iguanodon" – and referred specimens from Romania and elsewhere |journal=Revue de Paléobiologie |volume=28 |issue=1 |pages=211–273 |url=http://www.ville-ge.ch/mhng/paleo/paleo-pdf/28-1/pal-28-1-11.pdf | accessdate= 7 January 2010 }}
42. ^{{cite journal | last1 = Jin | first1 = L. | last2 = Chen | first2 = J. | last3 = Zan | first3 = S. | last4 = Godefroit | first4 = P. | year = 2009 | title = A new basal neoceratopsian dinosaur from the Middle Cretaceous of Jilin Province, China | url = | journal = Acta Geologica Sinica (English Edition) | volume = 83 | issue = 2| pages = 200–206 | doi = 10.1111/j.1755-6724.2009.00023.x }}
43. ^Jin, Chen, Zan, Godefroit.
44. ^Longrich, N.R. and Currie, P.J. (2009). "A microraptorine (Dinosauria–Dromaeosauridae) from the Late Cretaceous of North America." Proceedings of the National Academy of Sciences, 106(13) .
45. ^{{cite journal | last1 = You | first1 = H.-L. | last2 = Li | first2 = D.-Q. | year = 2009 | title = A new basal hadrosauriform dinosaur (Ornithischia: Iguanodontia) from the Early Cretaceous of northwestern China | url = | journal = Canadian Journal of Earth Sciences | volume = 46 | issue = 12| pages = 949–957 | doi = 10.1139/E09-067 |bibcode = 2009CaJES..46..949Y | last3 = Li | first3 = Da-Qing }}
46. ^Buffetaut, E., Suteethorn, V. and Tong, H. (2009). "An early 'ostrich dinosaur' (Theropoda: Ornithomimosauria) from the Early Cretaceous Sao Khua Formation of NE Thailand", pp. 229-243 IN E. Buffetaut, G. Cuny, J. Le Loeuff & V. Suteethorn (eds.), Late Palaeozoic and Mesozoic Ecosystems in SE Asia. Geological Society, London, Special Publications 315: 229-243. {{doi|10.1144/SP315.16}}
47. ^{{cite journal |first=A.H. |last=Turner |first2=S.J. |last2=Nesbitt |first3=M.A. |last3=Norell |title=A Large Alvarezsaurid from the Cretaceous of Mongolia |journal=American Museum Novitates |volume=3648 |pages=1–14 |year=2009 |doi=10.1206/639.1}}
48. ^F. Li; Peng G.; Ye Y.; Jiang S.; and Huang, D. (2009). "A new carnosaur from the Late Jurassic of Qianwei, Sichuan, China". Acta Geologica Sinica 83(9): 1203–1213. Abstract.
49. ^Li, Peng, Jiang, Huang.
50. ^{{cite journal | last1 = Hans-Dieter | first1 = Sues | last2 = Averianov | first2 = Alexander | year = 2009 | title = A new basal hadrosauroid dinosaur from the Late Cretaceous of Uzbekistan and the early radiation of duck-billed dinosaurs | journal = Proceedings of the Royal Society B | volume = 276 | issue = 1667| pages = 2549–2555 | doi = 10.1098/rspb.2009.0229 | pmid=19386651 | pmc=2686654}}
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53. ^Lü, Xu, Jiang, Jia, Li, Yuan, Zhang, Ji.
54. ^{{cite journal | last1 = González Riga | first1 = B. J. | last2 = Previtera | first2 = E. | last3 = Pirrone | first3 = C.A. | year = 2009 | title = Malarguesaurus florenciae gen. et sp. nov., a new titanosauriform (Dinosauria, Sauropoda) from the Upper Cretaceous of Mendoza, Argentina | url = | journal = Cretaceous Research | volume = 30 | issue = 1| pages = 135–148 | doi=10.1016/j.cretres.2008.06.006}}
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59. ^Sereno, Brusatte, Kriegstein, Zhao, Cloward.
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61. ^Lu, Xu, Jia, Zhang, Zhang, Yang, You, Ji.
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63. ^Brusatte, Benson, Chure, Xu, Sullivan, Hone.
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65. ^Wu, Currie, Dong, Pan, Tang.
66. ^{{cite journal | last1 = Ji | first1 = Q. | last2 = Ji | first2 = S.-A. | last3 = Zhang | first3 = L.-J. | year = 2009 | title = First large tyrannosauroid theropod from the Early Cretaceous Jehol Biota in northeastern China | url = | journal = Geological Bulletin of China | volume = 28 | issue = 10| pages = 1369–1374 }}
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103. ^{{Cite journal |author1=Gerald Mayr |author2=Nikita V. Zelenkov | year = 2009 | title = New Specimens of Zygodactylid Birds from the Middle Eocene of Messel, with Description of a New Species of Primozygodactylus | url=https://www.app.pan.pl/archive/published/app54/app54-015.pdf | journal = Acta Palaeontologica Polonica | volume = 54 | issue = 1 | pages = 15–20 | doi=10.4202/app.2009.b103}}
104. ^{{Cite journal|author=Eric M. Morschhauser |author2=David J. Varricchio |author3=Gao Chunling |author4=Liu Jinyuan |author5=Wang Xuri |author6=Cheng Xiadong |author7=Meng Qingjin|year=2009|title=Anatomy of the Early Cretaceous bird Rapaxavis pani, a new species from Liaoning Province, China |journal=Journal of Vertebrate Paleontology|volume=29|issue=2|pages=545–554|doi=10.1671/039.029.0210 }}
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107. ^{{Cite journal | author=Gerald Mayr | year = 2009 | title = A Small Loon and a New Species of Large Owl from the Rupelian of Belgium (Aves: Gaviiformes, Strigiformes) | url=http://www.senckenberg.de/files/content/forschung/abteilung/terrzool/ornithologie/belgium_owl.pdf | journal = Paläontologische Zeitschrift | volume = 83 | issue = 2 | pages = 247–254 | doi=10.1007/s12542-009-0019-3}}
108. ^{{Cite journal|author1=Jingmai K. O'Connor |author2=Xuri Wang |author3=Luis M. Chiappe |author4=Chunling Gao |author5=Qingjin Meng |author6=Xiaodong Cheng |author7=Jinyuan Liu |year=2009|title=Phylogenetic Support for a Specialized Clade of Cretaceous Enantiornithine Birds with Information from a New Species |url=http://geology.usc.edu/~jingmai/oconnoretal2009shanweiniao.pdf |journal=Journal of Vertebrate Paleontology|volume=29|issue=1|pages=188–204|doi=10.1080/02724634.2009.10010371}}
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111. ^{{cite journal |author1=Dalla Vecchia |author2=Fabio M. |year=2009 |title= Anatomy and systematics of the pterosaur Carniadactylus (gen. n.) rosenfeldi (Dalla Vecchia, 1995) |journal= Rivista Italiana de Paleontologia e Stratigrafia |volume= 115 |issue= 2 |pages=159–188}}
112. ^{{cite journal |doi= 10.1111/j.1755-6724.2009.00062.x |author= Lü, J. |year=2009 |title= A new non-pterodactyloid pterosaur from Qinglong County, Hebei Province of China |journal= Acta Geologica Sinica (English Edition) |volume= 83 |issue= 2 |pages=189–199}}
113. ^{{cite journal |doi= 10.1111/j.1755-6724.2009.00001.x |author= Lü J. |year=2009 |title= A baby pterodactyloid pterosaur from the Yixian Formation of Ningcheng, Inner Mongolia, China |journal= Acta Geologica Sinica |volume= 83 |issue= 1 |pages=1–8}}
114. ^{{cite journal |author1=Wang X. |author2=Kellner, A. W. A. |author3=Jiang S. |author4=Meng X. |year=2009 |title= An unusual long-tailed pterosaur with elongated neck from western Liaoning of China |journal= Anais da Academia Brasileira de Ciências |volume= 81 |issue= 4 |pages= 793–812 |doi= 10.1590/S0001-37652009000400016}}

References

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1 : 2009 in paleontology

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