1. Description

2. Taxonomy
     History    Pre-Darwinian    Post-Darwinian    {{anchor|Phylogenetic era}}Phylogenetic era    Subdivision    Angiosperm Phylogeny Group    Genera  

3. Distribution

4. Cultivation and uses

5. See also

6. References

7. Bibliography
     Books    Historical    Modern    Chapters    Symposia    Articles and theses    APG system    Pharmacology    Websites    Databases  

8. External links

• Agapanthoideae
• Allioideae
• Amaryllidoideae

The Amaryllidaceae are a family of herbaceous, mainly perennial and bulbous (rarely rhizomatous) flowering plants in the monocot order Asparagales. The family takes its name from the genus Amaryllis and is commonly known as the amaryllis family. The leaves are usually linear, and the flowers are usually bisexual and symmetrical, arranged in umbels on the stem. The petals and sepals are undifferentiated as tepals, which may be fused at the base into a floral tube. Some also display a corona. Allyl sulfide compounds produce the characteristic odour of the onion subfamily (Allioideae).

The family, which was originally created in 1805, now contains about 1600 species, divided into about 75 genera,^[1] 17 tribes and three subfamilies, the Agapanthoideae (agapanthus), Allioideae (onions and chives) and Amaryllidoideae (amaryllis, daffodils, snowdrops). Over time, it has seen much reorganisation and at various times was combined with the related Liliaceae. Since 2009, a very broad view has prevailed based on phylogenetics, and including a number of other former families.

The family is found in tropical to subtropical areas of the world and includes many ornamental garden plants and vegetables.

Description

The Amaryllidaceae are mainly terrestrial (rarely aquatic) flowering plants that are herbaceous or succulent geophytes (occasionally epiphytes) that are perennial, with the exception of four species. Most genera grow from bulbs, but a few such as Agapanthus, Clivia and Scadoxus develop from rhizomes (underground stems).{{sfn|Dimitri|1987}}

The leaves are simple rather fleshy and two-ranked with parallel veins. Leaf shape may be linear, strap like, oblong, elliptic, lanceolate (lance shaped) or filiform (threadlike). The leaves which are either grouped at the base or arranged alternatively on the stem may be sessile or petiolate and possess a meristem.

The flowers, which are hermaphroditic (bisexual), are actinomorphic (radially symmetrical), rarely zygomorphic, pedicellate or sessile, and are typically arranged in umbels at the apex of leafless flowering stems, or scapes and associated with a filiform (thread like) bract. The perianth (perigonium) consists of six undifferentiated tepals arranged in two whorls of three. The tepals are similar in shape and size, and may be free from each other or fused at the base (connate) to form a floral tube (hypanthium). In some genera, such as Narcissus, this may be surmounted by cup or trumpet shaped projection, the corona (paraperigonium or false corolla). This may be reduced to a mere disc in some species.

The position of the ovary varies by subfamily, the Agapanthoideae and Allioideae have superior ovaries, while the Amaryllidoideae have inferior ovaries. The six stamens are arranged in two whorls of three, occasionally more as in Gethyllis (Amaryllidoideae, 9–18).

The fruit is dry and capsule-shaped, or fleshy and berry-like.

The Allioideae produce allyl sulfide compounds which give them their characteristic smell.{{sfn|McGary|2001}}{{sfn|Rossi|1990}}

Taxonomy

History

Pre-Darwinian

Linnaeus described the type genus Amaryllis, from which the family derives its name, in his Species Plantarum in 1753,{{sfn|Linnaeus|1753|loc=[https://www.biodiversitylibrary.org/item/13829#page/304/mode/1up Amaryllis I pp. 292–293]}} with nine species, in the Hexandria monogynia (i.e. six stamens and one pistil){{sfn|Linnaeus Sexual System|2015}} containing 51 genera in all{{sfn|Linnaeus|1753|loc=[https://www.biodiversitylibrary.org/item/13829#page/297/mode/1up Hexandria monogynia I pp. 285–332]}} in his sexual classification scheme. The name Amaryllis had been applied to a number of plants over the course of history.

Hexandria monogynia has come to be treated as either liliaceous or amaryllidaceaous (see Taxonomy of Liliaceae) over time.{{sfn|Meerow et al.|1999}} From 1763, when Adanson conceived of these genera as 'Liliaceae'{{sfn|Adanson|1763|loc=[https://www.biodiversitylibrary.org/item/6958#page/588/mode/1up VIII. Liliaceae. Part II. p. 42]}} it was included in this family, placing Amaryllis in Section VII, Narcissi.{{sfn|Adanson|1763|loc=[https://www.biodiversitylibrary.org/item/6958#page/601/mode/1up VIII. Liliaceae Sectio VII. Part II. pp. 55–57]}} of his scheme, in which the Liliaceae had eight sections.

With de Jussieu came the formal establishment of organising genera into families (ordo) in 1789.{{Sfn|Jussieu|1789}} De Jussieu established the hierarchical system of taxonomy (phylogeny), placing Amaryllis and 15 related genera within a division of monocotyledons, a class (III) of Stamina Perigynia{{Sfn|Jussieu|1789|loc=[https://www.biodiversitylibrary.org/item/7125#page/130/mode/1up Stamina Perigynia p. 35]}} and 'order' Narcisse, divided into three subfamilies.{{Sfn|Jussieu|1789|loc=[https://www.biodiversitylibrary.org/item/7125#page/149/mode/1up Narcisse pp. 54–56]}} This system also formally described the Liliaceae, which were a separate order within the Stamina perigynia (Lilia). The use of the term Ordo (order) at that time was closer to what we now understand as family, rather than order.{{sfn|ICN|2011|loc=Names of families and subfamilies, tribes and subtribes p. 18.2}}{{sfn|Candolle|1813|loc=[https://www.biodiversitylibrary.org/item/88297#page/204/mode/1up Des familles et des tribus pp. 192–195]}} In creating his scheme, De Jussieu used a modified form of Linnaeus' sexual classification, but with the respective topography of stamens to carpels rather than just their numbers.

The Amaryllidaceae family was formally named as 'Amaryllidées' (Amaryllideae) in 1805, by Jean Henri Jaume Saint-Hilaire.{{sfn|Jaume Saint-Hilaire|1805|loc=[https://books.google.com/books?id=VEQAAAAAQAAJ&pg=RA1-PA130#v=onepage&q&f=false Amaryllidées vol. 1. pp. 134–142]}} In 1810 Brown proposed that a subgroup of Liliaceae be distinguished on the basis of the position of their ovaries (inferior) and be referred to as Amaryllideae{{sfn|Brown|1810|loc=[https://www.biodiversitylibrary.org/item/21871#page/164/mode/1up Prodromus. Amaryllideae p. 296]}} and in 1813 de Candolle described Liliacées Juss. and Amaryllidées Brown as two quite separate families.{{sfn|Candolle|1813|loc=[https://www.biodiversitylibrary.org/item/88297#page/231/mode/1up Esquisse. D'une Série linéaire et par conséquent artificielle, pour la disposition des familles naturelles du règne végetal p. 219]}} The literature on the organisation of genera into families and higher ranks became available in the English language with Samuel Frederick Gray's A natural arrangement of British plants (1821).{{sfn|Gray|1821}} Gray used a combination of Linnaeus' sexual classification and Jussieu's natural classification to group together a number of families having in common six equal stamens, a single style and a perianth that was simple and petaloid, but did not use formal names for these higher ranks. Within the grouping, he separated families by the characteristics of their fruit and seed. He treated groups of genera with these characteristics as separate families, such as Amaryllideae, Liliaceae, Asphodeleae, and Asparageae.{{sfn|Gray|1821|loc=[https://www.biodiversitylibrary.org/item/95185#page/10/mode/1up p.vi]}}

John Lindley (1830, 1846) was the other important British taxonomist of the early 19th century. In his first taxonomic work, An Introduction to the Natural System of Botany (1830){{sfn|Lindley|1830}} he partly followed De Jussieu by describing a subclass he called 'Endogenae, or Monocotyledonous Plants' (preserving de Candolle's Endogenæ phanerogamæ){{sfn|Lindley|1830|loc=[https://www.biodiversitylibrary.org/item/31944#page/319/mode/1up Endogenae, or Monocotyledonous Plants p. 251]}} divided into two tribes, the Petaloidea and Glumaceae. He divided the former, often referred to as petaloid monocots, into 32 orders, including the Amaryllideae.{{sfn|Lindley|1830|loc=[https://www.biodiversitylibrary.org/item/31944#page/327/mode/1up Amaryllideae The Narcissus Tribe p. 259]}} He defined the latter as "Hexapetaloideous bulbous hexandrous monocotyledons, with an inferior ovarium, a six-parted perianthium with equitant sepals, and flat, spongy seeds" and included Amaryllis, Phycella, Nerine, Vallota, and Calostemma.

By 1846, in his final scheme{{sfn|Lindley|1846}} Lindley had greatly expanded and refined the treatment of the monocots, introducing both an intermediate ranking (Alliances) and tribes within families. Lindley placed the Liliaceae within the Liliales, but saw it as a paraphyletic ("catch-all") family, being all Liliales not included in the other orders, but hoped that the future would reveal some characteristic that would group them better. This kept the Liliaceae{{sfn|Lindley|1846|loc=[https://www.biodiversitylibrary.org/item/32198#page/274/mode/1up Liliaceae - Lilyworts p. 200]}} separate from the Amaryllidaceae{{sfn|Lindley|1846|loc=[https://www.biodiversitylibrary.org/item/32198#page/229/mode/1up Amaryllidaceae - Amaryllids p. 155]}} (Narcissales Alliance). Of these Liliaceae{{sfn|Lindley|1846|loc=[https://www.biodiversitylibrary.org/item/32198#page/274/mode/1up Liliaceae - Lilyworts p. 200]}} was divided into eleven tribes (with 133 genera) and Amaryllidaceae{{sfn|Lindley|1846|loc=[https://www.biodiversitylibrary.org/item/32198#page/229/mode/1up Amaryllidaceae - Amaryllids p. 155]}} into four tribes (with 68 genera), yet both contained many genera that would eventually segregate to each other's contemporary orders (Liliales and Asparagales respectively). The Liliaceae would be reduced to a small 'core' represented by the Tulipeae tribe (18 genera), while large groups such Scilleae and Asparagae would become part of Asparagales either as part of the Amaryllidaceae or as separate families. While of the four tribes of the Amaryllidaceae, the Amaryllideae and Narcissea would remain as core amaryllids while the Agaveae would be part of Asparagaceae, but the Alstroemeriae would become a family within the Liliales.

Since then, seven of Linnaeus' Hexandria monogynia genera have consistently been placed in a common taxonomic unit of amaryllids, based on the inferior position of the ovaries (whether this be as an order, suborder, family, subfamily, tribe or section).{{sfn|Meerow et al.|2000a}} Thus, much of what we now consider Amaryllidaceae remained in Liliaceae because the ovary was superior, till 1926 when John Hutchinson transferred them to Amaryllidaceae.{{sfn|Hutchinson|1926}} This usage of the family entered the English language literature through the work of Samuel Frederick Gray (1821),{{sfn|Gray|1821|loc=[https://www.biodiversitylibrary.org/item/95185#page/202/mode/1up Fam. XIV Amaryllideae Brown. II p. 190–193; Liliaceae p. 173]}} William Herbert (1837){{sfn|Herbert|1837}} and John Lindley (1830,{{sfn|Lindley|1830|loc=[https://www.biodiversitylibrary.org/item/31944#page/327/mode/1up CCXXXVIII Amaryllideae The Narcissus Tribe pp. 259–260; Liliaceae p. 279]}} 1846{{sfn|Lindley|1846|loc=[https://www.biodiversitylibrary.org/item/32198#page/229/mode/1up Order XLVI Amaryllidaceae—Amaryllids pp. 155–158; Liliaceae p. 200 ]}}). Meanwhile, Lindley had described two Chilean genera which for which he created a new family, Gilliesieae.{{sfn|Lindley|1846|loc=[https://www.biodiversitylibrary.org/item/31944#page/343/mode/1up CCXLVIII Gilliesieae. pp. 275-277]}}

The number of known genera within these families continued to grow, and by the time of the Bentham and Hooker classification (1883), the Amaryllidaceae (Amaryllideae) were divided into four tribes, of which only one (Amarylleae) is still included.{{sfn|Bentham|Hooker|1883|loc=[https://www.biodiversitylibrary.org/item/14690#page/272/mode/1up Vol. 3, Part 2. Amaryllideae pp. 711–740]}} The Liliaceae{{sfn|Bentham|Hooker|1883|loc=[https://www.biodiversitylibrary.org/item/14690#page/309/mode/1up Vol. 3, Part 2. Liliaceae pp. 748–836]}} were becoming one of the largest families, and Bentham and Hooker divided it into 20 tribes, of which one was the Allieae,{{sfn|Bentham|Hooker|1883|loc=[https://www.biodiversitylibrary.org/item/14690#page/359/mode/1up Vol. 3, Part 2. Allieae pp. 798–807]}} which as Allioideae would eventually become part of Amaryllidaceae as two of its three subfamilies. The Allieae included both Agapantheae,{{sfn|Bentham|Hooker|1883|loc=[https://www.biodiversitylibrary.org/item/14690#page/359/mode/1up Vol. 3, Part 2. Agapantheae p. 798]}} the third of the current subfamilies, and Lindley's Gilliesieae{{sfn|Bentham|Hooker|1883|loc=[https://www.biodiversitylibrary.org/item/14690#page/365/mode/1up Vol. 3, Part 2. Gilliesieae pp. 804–806]}} as two of its four subtribes.{{sfn|Bentham|Hooker|1883|loc=[https://www.biodiversitylibrary.org/item/14690#page/311/mode/1up Vol. 3, Part 2. Allieae (Conspectus) pp. 750]}} Bentham and Hooker's scheme was the last major classification using the natural approach.{{sfn|Stuessy|2009|loc=[https://books.google.com/books?id=b9Q2EOkw7toC&pg=PA47 Natural classification p. 47]}}

Post-Darwinian

Although Charles Darwin's Origin of Species (1859) preceded Bentham and Hooker's publication, the latter project was commenced much earlier and Bentham was initially sceptical of Darwinism.{{sfn|Stuessy|2009|loc=[https://books.google.com/books?id=b9Q2EOkw7toC&pg=PA47 Natural classification p. 47]}} The new phyletic approach changed the way that taxonomists considered plant classification, incorporating evolutionary information into their schemata. The major works in the late 19th and early 20th centuries employing this approach were German, those of Eichler (1875–1886), Engler, Prantl (1886–1924), and Wettstein (1901–1935).

The Amaryllidaceae were treated similarly in the German-language literature to the manner they had been in English. August Eichler (1886){{sfn|Eichler|1886}} was the first phyletic taxonomist and positioned the Amaryllidaceae and Liliaceae within the Liliiflorae,{{sfn|Eichler|1886|loc=[https://books.google.com/books?id=XE0bAAAAYAAJ&pg=PA34 Liliiflorae p. 34]}}{{sfn|Wettstein|1924|loc=Liliiflorae p. 862}} one of the seven orders of monocotyledons. Liliaceae included both Allium and Ornithogalum (modern Allioideae). Adolf Engler developed Eichler's ideas much further, into much more elaborate schemes that evolved over time, from his 1888 scheme, contributed by Pax{{sfn|Pax|1888}} to his 1903 version{{sfn|Engler|1903|loc=[https://www.biodiversitylibrary.org/item/63778#page/127/mode/1up Liliiflorae p. 93]}} In the latter, the Liliineae were a suborder of Liliiflorae, including both Liliaceae and Amaryllidaceae families. Within the Liliaceae, the core liliids were segregated in subfamily Lilioideae from the alliaceous subfamily, Allioideae. Allieae, Agapantheae, and Gilliesieae were the three tribes within this subfamily.{{sfn|Engler|1903|loc=[https://www.biodiversitylibrary.org/item/63778#page/130/mode/1up Subfamily Allioideae p. 96]}} A somewhat similar approach to Liliiflorae{{sfn|Wettstein|1924|loc=Liliiflorae p. 862}} was adopted by Wettstein (without suborders or tribes), and with Alliodeae (Allium) and Lilioideae (Ornithogalum) as subfamilies of Liliaceae.{{sfn|Wettstein|1924|loc=Liliaceae p. 863}} Wettstein's Amaryllidaceae contained three subfamilies.,{{sfn|Wettstein|1924|loc=Amaryllidaceae p. 871}} including Amaryllidoideae and Agavoideae.

The early 20th century was marked by increasing doubts about the placement of the alliaceous genera within Liliaceae. Lotsy was the first taxonomist to propose separating them, and in his system he describes Agapanthaceae, Alliaceae, and Gilliesiaceae as new and separate families from Liliaceae.{{sfn|Lotsy|1911|loc=[https://www.biodiversitylibrary.org/item/77531#page/740/mode/1up Agapanthaceae, Alliaceae, Gilliesiaceae pp. 732–734]}} This approach was adopted by a number of other authorities, such as Dahlgren (1985){{sfn|Dahlgren|Clifford|Yeo|1985|loc=[https://books.google.com/books?id=3iGndTFY0skC&pg=PA193 Alliaceae pp. 193–198]}} and Rahn (1998).{{sfn|Kubitzki|1998|loc=[https://link.springer.com/chapter/10.1007%2F978-3-662-03533-7_9 K. Rahn. Alliaceae pp. 70–78]}}

Another approach was that of John Hutchinson (1926), who performed the first major recircumscription of the family in over a century. He doubted Brown's dictum that the position of the ovary was the distinguishing feature that separated Amaryllidaceae and Liliaceae. He treated Amaryllidaceae as bulbous plants with umbellate inflorescences, the latter characteristic being the defining feature: "an umbellate inflorescence subtended by an involucre of one or more spathaceous bracts".{{sfn|Hutchinson|1926}} His work on this has been upheld by subsequent research and his definition remains valid today.{{sfn|Wilkin|2012}} Using this criterion, he removed a number of taxa (Agavaceae, Hypoxidaceae, Alstroemeriaceae) and transferred the Agapantheae, Allieae, and Gilliesieae from Liliaceae to Amaryllidaceae.{{sfn|Hutchinson|1926}}

Other writers proposed reuniting Amaryllidaceae with Liliaceae. Thorne (1976){{sfn|Thorne|1976}} and Cronquist (1988){{sfn|Cronquist|1988}} both included Amaryllidaceae within a broad concept of Liliaceae{{sfn|Meerow et al.|2000a}} (although Thorne later separated them again, but keep Alliaceae as a third family).{{sfn|Thorne|1992}} Thus 'Alliaceae' were variously included in either Liliaceae, Amaryllidaceae, or as a separate entity. This uncertainty of circumscription reflected a wider problem with the petaloid monocots in general. Over the course of time, widely differing views as to the limits of the family have been expressed, so much of the literature dealing with this family requires careful inspection to determine which sense of the Amaryllidaceae the work treats.

{{anchor|Phylogenetic era}}Phylogenetic era

The current phylogenetic era of understanding the taxonomic relationships of Amaryllidaceae began with the work of Fay and Chase (1996) who used the plastid gene rubisco rbcL to identify the close relationship between Agapanthus, Alliaceae, and Amaryllidaceae.{{sfn|Fay|Chase|1996}} Agapanthus had variously been included in Alliaceae or was placed in a separate family, Agapanthaceae. They relocated Agapanthus within Amaryllidaceae which they considered a sister group to Amaryllidaceae. Nevertheless, the Angiosperm Phylogeny Group (APG) classification (1998) still considered these three separate families within Asparagales.{{sfn|APG|1998}} The close relationship was confirmed in a more detailed study by Meerow (1999) who confirmed the monophyly of Amaryllidaceae, with Agapanthaceae as its sister family and Alliaceae in turn as sister to the Amaryllidaceae/Agapanthaceae clade.{{sfn|Meerow et al.|1999}}

In its second iteration (2003), the APG proposed simplifying the higher (core) Asparagales by reducing them to two more broadly circumscribed families, and provisionally proposed the name Alliaceae sensu lato (s.l.) to include the three sister families (Agapanthaceae, Alliaceae sensu stricto, s.s., and Amaryllidaceae), since together they form a monophyletic group. In this respect, they were following Hutchinson's system (see above). Under this proposal, the three families became reduced to subfamilies (and by extension the subfamilies of Alliaceae s.s. being reduced to tribes.) At the same time, they appreciated an argument exisyed for making Amaryllidaceae s.l. the formal name of the new and larger family,{{sfn|APG|2003}}{{sfn|Kamenetsky|2012|loc=[https://books.google.com/books?id=K0LNBQAAQBAJ&lpg=PA25 p. 25]}} a position subsequently strongly supported by Meerow and colleagues.{{sfn|Meerow|Snijman|2006}}{{sfn|Meerow et al.|2007}}

The 2009 version of the APG formally adopted this broad view and the conserved name Amaryllidaceae. To distinguish this broader family from the older, narrower family, it has become customary to refer to Amaryllidaceae sensu APG, or as used by APG, Amaryllidaceae s.l.. as opposed to Amaryllidaceae s.s..{{sfn|APG|2009}}{{sfn|Chase et al.|2009}}

This phylogenetic tree (cladogram) shows the placement of Amaryllidaceae s.l. within the order Asparagales.{{sfn|ps=none|Stevens|2016|loc=Amaryllidaceae}}

             }}           }}         }}       }}

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Subdivision

As reconstituted by the APG, Amaryllidaceae s.l. consists of three subfamilies, Agapanthoideae, Allioideae, and Amaryllidoideae, corresponding to the three families that were subsumed into it:{{sfn|Garcia et al|2014}}

• Agapanthoideae (Agapanthaceae)
• Allioideae (Alliaceae)
• Amaryllidoideae (Amaryllidaceae s.s.)

Of these, one (Agapanthoideae) is monogeneric for Agapanthus (see Cladogram I).

Of the other two subfamilies, Allioideae was resolved into three subdivisions by the initial phylogenetic studies of Fay and Chase (1996). Since they treated Allioideae as family Alliaceae, these were subfamilies Allioideae, Tulbaghioideae, and Gilliesioideae. When family Alliaceae was reduced to subfamily Allioideae, they were reduced to tribes, namely Allieae, Tulbaghieae and Gilliesieae (see Cladogram II).{{sfn|Fay|Chase|1996}}

             }}      }}

Complete resolution of infrafamilial (suprageneric) relationships within subfamily Amaryllidoideae (Amaryllidaceae s.s.) has proven more difficult.{{sfn|Meerow|Snijman|2006}} Fay and Chase's study lacked sufficient resolution for further elucidation of this group. Historically a wide variety of infrafamilial classification systems have been proposed for the Amaryllidaceae. In the latter twentieth century there were at least six schemes, including Hutchinson (1926),{{sfn|Hutchinson|1926}} Traub (1963),{{sfn

The further application of molecular phylogenetics produced a complex picture that only partially related to the tribal structure considered up to that date, which had been based on morphology alone.{{sfn|Meerow et al.|1999}} RAther Amaryllidaceae resolved along biogeographical lines. A predominantly South African clade identified as Amaryllideae was a sister group to the rest of the family. The two other African tribes were Haemantheae and Cyrtantheae, and an Australasian tribe Calostemmateae was also identified, but a large clade could only be described as Eurasian and American, each of which were monophyletic sister clades to each other. The Eurasian clade was poorly resolved with the exception of Lycorideae (Central and East Asian). The American clade was better resolved identifying both Hippeastreae as a tribe (and Zephyranthinae as a subtribe within it). The American clade also included an Andean clade{{sfn|Meerow et al.|1999}}

Further investigation of the American clade suggested the presence of two groups, the Andean clade and a further "Hippeastroid" clade, in which Griffineae was sister to the rest of the clade (Hippeastreae). Similarly within the Andean clade Eustephieae appeared as sister to the remaining clade, including Hymenocallideae. A new tribe, Clinantheae was also identified in this group.{{sfn|Meerow et al.|2000b}}

The Eurasian clade was also further resolved (for historical treatment, see Table I Meerow et al. 2006) into four tribes, Pancratieae, Narcisseae, Galantheae and Lycorideae. This positioned Lycorideae as sister to the remaining Mediterranean tribes.{{sfn|Meerow et al|2006a}}

These relationships are summarised in the following cladogram:

Angiosperm Phylogeny Group

Publication of the third version of the APG classification and acceptance of Amaryllidaceae s.l.{{sfn|APG|2009}} was accompanied by a listing of accepted subfamily and tribal names, since the change in rank from family to subfamily necessitated a revision of other lower ranks, as follows:{{sfn|Chase et al.|2009}}

Family: Amaryllidaceae J.St.-Hil., Expos. Fam. Nat. 1: 134. Feb–Apr 1805, nom. cons.

• Subfamily: Agapanthoideae Endl., Gen. Pl.: 141. Dec 1836.
• Subfamily: Allioideae Herb., Amaryllidaceae: 48. late Apr 1837.
  • Tribe Allieae Dumort., Fl. Belg.: 139. 1827. (3 tribes)
  • Tribe Gilliesieae Baker, J. Linn. Soc., Bot. 14: 509. 24 Apr 1875. (18 genera)
  • Tribe Tulbaghieae Endl. ex Meisn., Pl. Vasc. Gen.: Tab. Diagn. 397, 399, Comm. 302. 17–20 Dec 1842.
• Subfamily: Amaryllidoideae Burnett, Outl. Bot.: 446. Feb 1835 (15 tribes)
  • Tribe Amaryllideae Dumort., Anal. Fam. Pl.: 58. 1829.
  • Tribe Calostemmateae D.Müll.-Doblies & U.Müll.Doblies, Feddes Repert. 107 (Short commun.): 7 Dec 1996.
  • Tribe Cyrtantheae Traub, Herbertia 5: 111. Nov 1938.
  • Tribe Eucharideae Hutch., Fam.Fl.Pl.2:130.20 Jul 1934.
  • Tribe Eustephieae Hutch., Fam.Fl.Pl.2:130.20 Jul 1934.
  • Tribe Galantheae Parl., Fl. Ital. 3: 75. 1858.
  • Tribe Gethyllideae Dumort., Anal. Fam. Pl.: 58. 1829.
  • Tribe Haemantheae Hutch., Fam. Fl. Pl. 2: 130. 20 Jul 1934.
  • Tribe Hippeastreae Herb. ex Sweet, Brit. Fl. Gard., ser. 2, 1: ad t. 14. 1 Sep 1829.
  • Tribe Hymenocallideae Small, Man. S.E. Fl.: 315. 30 Nov 1933.
  • Tribe Lycorideae Traub ex D.Müll.-Doblies & U.Müll.Doblies, Feddes Repert. 107 (Short commun.): 6. Dec. 1996.
  • Tribe Narcisseae Lam. & DC., Syn. Pl. Fl. Gall.: 165. 30 Jun 1806.
  • Tribe Pancratieae Dumort., Anal. Fam. Pl.: 58. 1829.
  • Tribe Stenomesseae Traub, Pl. Life 19: 60. Jan 1963

This circumscription differs from the phylogenetic descriptions of Meerow and colleagues in several respects. Griffineae is recognised as a distinct tribe within the Hippeastroid clade, and Stenomesseae is recognised as polyphyletic with two distinct types based on leaf shape (lorate-leafed and petiolate-leafed). The lorate-leafed species of the type genus of Stenomesseae, Stemomesson, were transferred to a new tribe, Clinantheae as sister to Hymenocallideae in the Andean clade. The remnants of Stemomesson then formed a distinct clade with Eucharis (Eucharidae) and Eucharidae renamed as Stenomesseae (see Cladogram III).{{sfn|Weber|Wilkin|2007}}{{sfn|Meerow et al|2004}}{{sfn|Meerow et al.|2000b}}{{sfn|PBS|2012|loc=Clinanthus}}{{sfn|Byng|2014|loc=[https://books.google.com/books?id=yoLaBAAAQBAJ&pg=PA84 Amaryllidaceae]}}

• Tribe Griffineae Ravenna
• Tribe Clinantheae Meerow

Genera

The Angiosperm Phylogeny Website lists 73 genera and 1,605 species within Amaryllidaceae s.l.,{{sfn|ps=none|Stevens|2016|loc=Amaryllidaceae}} while The Plant List (2013) gives 80 genera and 2,258 species.{{sfn|The Plant List|2013}}

Distribution

Amaryllidaceae are a cosmopolitan family, whose distribution is pantropical to subtropical, but infrafamilial relationships are related to geographical considerations. The Amaryllideae tribe is primarily South African, and Haemantheae and Cyrtantheae are also African, while the Calostemmateae are Australasian. Other elements are Eurasian and American, including an Andean subclade without necessarily following strictly tribal delimitations. This leads to discussions of, for instance American Amaryllidaceae.{{sfn|Meerow et al.|1999}}{{sfn|Meerow et al.|2000b}} The Eurasian clade includes Lycorideae. The American clade includes the Hippeastreae, Eustephieae and Zephyranthinae.{{sfn|Meerow et al.|1999}}

Cultivation and uses

The Amaryllidaceae include many ornamental garden plants such as daffodils, snowdrops and snowflake, pot plants such as amaryllis and Clivia, and vegetables, such as onions, chives, leeks and garlic. A number of tropical lily-like plants are also sold, such as the belladonna lily, Amazon lily, blood lily (Cape tulip), Cornish lily (Nerine), and the Eurasian winter daffodil, Sternbergia.

Their economic importance lies in floriculture for cut flowers and bulbs, and commercial vegetable production.

See also

• {{Portal-inline|size=tiny|Amaryllidaceae}}

References

Bibliography

Books

Historical

• {{cite book |url=https://www.biodiversitylibrary.org/bibliography/669#/summary |last=Linnaeus |first=C. |title=Species Plantarum: exhibentes plantas rite cognitas, ad genera relatas, cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas|year=1753 |publisher=Impensis Laurentii Salvii|location=Stockholm|accessdate=26 January 2015|authorlink=Linnaeus|ref=harv}} see also Species Plantarum
• {{cite book|last=Adanson|first=Michel|authorlink=Adanson|title=Familles des plantes|publisher=Vincent|location=Paris|url=https://www.biodiversitylibrary.org/bibliography/271#/summary|accessdate=9 February 2014|year=1763|ref=harv}}

[https://www.biodiversitylibrary.org/item/6958#page/521/mode/1up Table of 58 families, Part II: Page 1]

[https://www.biodiversitylibrary.org/item/6958#page/528/mode/1up Table of 1615 genera, Part II: Page 8]

• {{cite book |last=Jussieu |first=Antoine Laurent de |authorlink=Antoine Laurent de Jussieu|title=Genera Plantarum, secundum ordines naturales disposita juxta methodum in Horto Regio Parisiensi exaratam |year=1789 |location=Paris |url=https://www.biodiversitylibrary.org/bibliography/284| accessdate=26 January 2015|ref=harv|publisher=apud viduam Herissant et Theophilum Barrois }}
• {{cite book|last1=Jaume Saint-Hilaire|first1=Jean Henri|authorlink=Jaume Saint-Hilaire|title=Exposition de familles naturales|date=1805|publisher=Treutel et Würtz|location=Paris|url=https://books.google.com/books?id=VEQAAAAAQAAJ|accessdate=25 October 2014|ref=harv}}
• {{cite book|last1=Brown|first1=Robert|title=Prodromus florae Novae Hollandiae et Insulae Van-Diemen, exhibens characteres plantarum|date=1810|publisher=Taylor|location=London|url=https://www.biodiversitylibrary.org/bibliography/3678#/summary|accessdate=30 October 2014|ref=harv}}
• {{cite book|last1 = Candolle|first1=A. P. de|year = 1813|authorlink=A. P. de Candolle|title = Théorie élémentaire de la botanique, ou exposition des principes de la classification naturelle et de l'art de décrire et d'etudier les végétaux|url=https://www.biodiversitylibrary.org/bibliography/39705#/summary|accessdate=5 February 2014|language=french|ref=harv}}
• {{cite book|last=Gray|first=Samuel Frederick|authorlink=Samuel Frederick Gray|title=A natural arrangement of British plants: according to their relations to each other as pointed out by Jussieu, De Candolle, Brown, &c. including those cultivated for use; with an introduction to botany, in which the terms newly introduced are explained|year=1821|publisher=Baldwin|location=London|url=https://www.biodiversitylibrary.org/bibliography/43804#/summary|accessdate=2 February 2014|ref=harv}}
• {{cite book|last=Lindley|first=John|authorlink=John Lindley|title=An introduction to the natural system of botany : or, A systematic view of the organisation, natural affinities, and geographical distribution, of the whole vegetable kingdom : together with the uses of the most important species in medicine, the arts, and rural or domestic economy|year=1830|publisher=Longman|location=London|url=https://www.biodiversitylibrary.org/bibliography/7551|accessdate=2 February 2014|ref=harv}}
• {{cite book|last1=Herbert|first1=William|authorlink=William Herbert (botanist)|title=Amaryllidaceae: Preceded by an Attempt to Arrange the Monocotyledonous Orders, and Followed by a Treatise on Cross-bred Vegetables, and Supplement|url=https://books.google.com/books?id=dUcaAAAAYAAJ|accessdate = 26 January 2015|date=1837|publisher=Ridgway|location=London|ref=harv}}
• {{cite book|last=Lindley|first=John|authorlink=John Lindley|title=The Vegetable Kingdom: or, The structure, classification, and uses of plants, illustrated upon the natural system|publisher=Bradbury|location=London|accessdate=26 January 2015|year=1846| url = https://www.biodiversitylibrary.org/bibliography/7756#/summary|ref=harv}}
• {{cite book|last=Bentham|first=G.|last2= Hooker|first2=J.D.| authorlink1=George Bentham|authorlink2=Joseph Dalton Hooker|title=Genera plantarum ad exemplaria imprimis in herbariis kewensibus servata definita|format=3 vols. 1865–1883 |year=1883|publisher=L Reeve & Co.|location=London|url=https://www.biodiversitylibrary.org/bibliography/747#/summary|accessdate=24 January 2014|ref=harv}}
• {{cite book|last1=Eichler|first1=August W.|authorlink = August W. Eichler|title=Syllabus der Vorlesungen über specielle und medicinisch-pharmaceutische Botanik|date=1886|origyear=1876|publisher=Borntraeger|location=Berlin|edition=4th|url=https://books.google.com/books?id=XE0bAAAAYAAJ|ref=harv}}
• {{cite book|last=Baker|first=John Gilbert|authorlink=John Gilbert Baker|title=Handbook of the Amaryllideæ including the Alstrœmerieæ and Agaveæ|year=1888|publisher=Bell|location=London|url=https://www.biodiversitylibrary.org/bibliography/15516#/summary|ref=harv}}
• {{cite book|editor1-last=Engler|editor1-first=Adolf|editor2-last=Prantl|editor2-first=Karl|editorlink1=Adolf Engler|title=Die Natürlichen Pflanzenfamilien nebst ihren Gattungen und wichtigeren Arten, insbesondere den Nutzpflanzen, unter Mitwirkung zahlreicher hervorragender Fachgelehrten 1887–1915 II(5)|date=1888|publisher=W. Engelmann|location=Leipzig|url=https://www.biodiversitylibrary.org/bibliography/4635#/summary|accessdate=6 April 2015|ref=harv}}
• {{cite book|title=Das Pflanzenreich :regni vegetablilis conspectus|publisher=Engelmann|location=Leipzig|url=https://www.biodiversitylibrary.org/bibliography/250#/summary|editor-last= Engler|editor-first=Adolf|editorlink=Adolf Engler|accessdate=5 February 2014|year=1900–1968|ref=harv}}
• {{cite book|title=Syllabus der Pflanzenfamilien: eine Übersicht über das gesamte Pflanzensystem mit Berücksichtigung der Medicinal- und Nutzpflanzen nebst einer Übersicht über die Florenreiche und Florengebiete der Erde zum Gebrauch bei Vorlesungen und Studien über specielle und medicinisch-pharmaceutische Botanik|date=1903|publisher=Gebrüder Borntraeger Verlag|location=Berlin|url=https://www.biodiversitylibrary.org/bibliography/22956#/summary|edition=3|editor-last=Engler|editor-first=Adolf|editorlink=Adolf Engler|accessdate=31 January 2014|ref=harv}}
• {{cite book| first= Richard|last= Wettstein|authorlink=Richard Wettstein|year = 1924|edition=3rd|title = Handbuch der Systematischen Botanik 2 vols|url=http://biolib.mpipz.mpg.de/library/authors/author_00267_de.html|accessdate=15 April 2015|ref=harv}}

Modern

• {{cite book|last1=Byng|first1=James W.|title=The Flowering Plants Handbook: A Practical Guide to Families and Genera of the World|date=2014|publisher=Plant Gateway Ltd|isbn=978-0992999308|url=https://books.google.com/books?id=yoLaBAAAQBAJ|ref=harv}}
• {{cite book|last1=Lotsy|first1=J. P.|title=Vorträge über botanische stammesgeschichte, gehalten an der Reichsuniversität zu Leiden. Ein lehrbuch der pflanzensystematick. III Cormophyta Siphonogamia|date=1911|publisher=G. Fischer|location=Jena|url=https://www.biodiversitylibrary.org/bibliography/33103#/summary|accessdate=28 January 2015|ref=harv}}
• {{cite book|last=Hutchinson|first=John|authorlink=John Hutchinson (botanist)|title=The families of flowering plants, arranged according to a new system based on their probable phylogeny. 2 vols|date=1934|edition=1st|publisher=Macmillan|ref={{harvid|Hutchinson|1926}}}} Volume 1: Monocotyledonae 1926, Volume 2: Dicotyledonae 1934.
• {{cite book|last=Hutchinson|first=John|title=The families of flowering plants, arranged according to a new system based on their probable phylogeny. 2 vols|date=1959|edition=2|publisher=Macmillan|ref=harv}}
• {{cite book|last=Hutchinson|first=John|title=The families of flowering plants, arranged according to a new system based on their probable phylogeny. 2 vols|date=1973|edition=3|publisher=Oxford University Press|ref=harv}}
• {{cite book |last=Cronquist |first=A |title=An integrated system of classification of flowering plants |year=1981 |publisher=Columbia University Press |location=New York |url=https://books.google.com/books?id=EB6qflbmcrwC|isbn=9780231038805|ref=harv }}
• {{cite book|last1=Cronquist|first1=Arthur|title=The evolution and classification of flowering plants|date=1988|publisher=New York Botanical Garden|location=Bronx, New York|isbn=978-0893273323|edition=2|ref=harv}}
• {{cite book |last=Dahlgren |first=R.M. |last2=Clifford |first2=H.T. |last3=Yeo |first3=P.F. |title=The families of the monocotyledons |year=1985 |publisher=Springer-Verlag |location=Berlin|url=https://books.google.com/books?id=3iGndTFY0skC|isbn= 978-3-642-64903-5| accessdate=10 February 2014|ref=harv}}
• {{cite book|editor-last=Kubitzki|editor-first=K.|editorlink=Klaus Kubitzki|title=The families and genera of vascular plants. Vol.3|year=1998|publisher=Springer-Verlag|location=Berlin, Germany|isbn=978-3-540-64060-8|url=https://books.google.com/books?id=FyPVYzL76sMC|accessdate=14 January 2014|ref=harv}}
  • [https://link.springer.com/chapter/10.1007%2F978-3-662-03533-7_9 Rahn K. Alliaceae. pp. 70–78]
• {{cite book|last1=Rossi|first1=Rosella|title=Guía de bulbos|date=1990|publisher=Grijalbo|location=Barcelona|isbn=978-8425321658|ref=harv}}
• {{cite book|title=Ornamental Geophytes: From Basic Science to Sustainable Production|year=2012|publisher=CRC Press|isbn=978-1-4398-4924-8|url=https://books.google.com/books?id=5B-ucdbgA3wC|editor1=Rina Kamenetsky |editor2=Hiroshi Okubo |ref={{harvid|Kamenetsky|2012}}}}
  • [https://www.academia.edu/6359004/Meerow_-_Ch_2_Ornamental_geophytes_Taxonomy_and_Phylogeny Chapter 2. Alan Meerow. Taxonomy and Phylogeny. pp. 17–55]
• {{cite book|last1=Dimitri|first1=M|title=Enciclopedia Argentina de Agricultura y Jardinería. Tomo I. Descripción de plantas cultivadas|date=1987|publisher=Editorial ACME S.A.C.I.|location=Buenos Aires|ref=harv}} see Enciclopedia Argentina de Agricultura y Jardinería
• {{cite book|last1=McGary|first1=Mary Jane|title=Bulbs of North America|date=2001|publisher=Timber Press|isbn=978-0-88192-511-1|ref=harv}}
• {{cite book|last1=Hickey|first1=Michael|last2=King|first2=Clive|title=Common families of flowering plants|date=1997|publisher=Cambridge University Press|location=Cambridge|isbn=978-0521576093|url=https://books.google.com/books?id=nhQQho64fxYC|accessdate=6 February 2015|ref=harv}} At C.U.P.
• {{cite book|last=Stuessy|first=Tod F.|title=Plant Taxonomy: The Systematic Evaluation of Comparative Data|publisher=Columbia University Press|isbn=978-0-231-14712-5|url=https://books.google.com/books?id=0bYs8F0Mb9gC|accessdate=6 February 2014|year=2009|ref=harv}}
• {{cite book |last=Takhtadzhi︠a︡n |first=Armen Leonovich |authorlink=Takhtadzhi︠a︡n|title=Flowering Plants |year=2009 |publisher=Springer |location=New York|isbn=978-1-4020-9609-9 |url=https://books.google.com/books?id=oumyfO-NHuUC|accessdate=7 January 2014|ref=harv}}
• {{cite book|last1=Traub|first1=H. P.|authorlink=Hamilton Traub|title=Genera of the Amaryllidaceae|url=http://babel.hathitrust.org/cgi/pt?id=mdp.39015006929270;view=1up;seq=3|date=1963|publisher=American Plant Life Society|location=La Jolla, California|ref=harv}}

Chapters

• {{cite book|last1=Meerow|first1=AW|title=Flowering Plants · Monocotyledons|last2=Snijman|first2=DA|authorlink1=Alan Meerow|pages=83–110|doi=10.1007/978-3-662-03533-7_11|ref={{harvid|Meerow|Snijman|1998}}|year=1998|isbn=978-3-642-08377-8}}, in {{harvtxt|Kubitzki|1998}}. [https://books.google.com/books?id=FyPVYzL76sMC&pg=PA85 (additional excerpts)]
• {{cite book| last1 =Meerow| first1 =Alan W.|last2=Fay|first2= Michael F.|last3 = Chase|first3=Mark W.|last4 = Guy|first4=Charles L.|last5 = Li|first5 = Qin-Bao|last6 = Snijman|first6 = Deirdre|last7= Yang|first7 = Si-Lin |pages =372–386 | title= Phylogeny of Amaryllidaceae: Molecules and morphology | url=https://books.google.com/books?id=YzQBUQqLS0YC&pg=PA372|accessdate=25 January 2015| date=2000|isbn =9780643099296|ref={{harvid|Meerow et al.|2000a}}}}, in {{Harvtxt|Wilson|Morrison|2000}}
• {{cite book|last1=Meerow|first1=Alan|authorlink=Alan Meerow|title=Towards a phylogeny of the Amaryllidaceae|pages=169–179}}, in {{harvtxt|Rudall|Cribb|Cutler|Humphries|1995 }}
• {{cite book|last1=Pax|first1=Ferdinand|authorlink=Ferdinand Pax|title=Amaryllidaceae|volume=Teil 2, Abt. 1-6|url=https://www.biodiversitylibrary.org/item/56456#page/759/mode/1up|pages=97–124|ref={{harvid|Pax|1888}}|publisher=W. Engelmann}}, in {{harvtxt|Engler|Prantl|1888}}

Symposia

• {{cite book |editor1-last=Rudall |editor1-first=P.J. |editor2-last=Cribb |editor2-first=P.J. |editor3-last=Cutler |editor3-first=D.F. |editor4-last=Humphries |editor4-first=C.J. |editorlink1=Paula Rudall|year=1995 |title=Monocotyledons: systematics and evolution (Proceedings of the International Symposium on Monocotyledons: Systematics and Evolution, Kew 1993)|publisher=Royal Botanic Gardens |location=Kew |isbn=978-0-947643-85-0 |url=http://www.press.uchicago.edu/ucp/books/book/distributed/M/bo9856357.html |accessdate=14 January 2014 |ref=harv }}
• {{cite book|editor1-last= Wilson|editor1-first= K. L. |editor2-last=Morrison |editor2-first=D. A.|title=Monocots: Systematics and evolution (Proceedings of the Second International Conference on the Comparative Biology of the Monocotyledons, Sydney, Australia 1998)|date= 19 May 2000|publisher=CSIRO|location=Collingwood, Australia|url=http://www.publish.csiro.au/pid/2424.htm|isbn=978-0-643-06437-9|accessdate=14 January 2014 |ref={{harvid|Wilson|Morrison|2000}}}} [https://books.google.com/books?id=YzQBUQqLS0YC Excerpts]

Articles and theses

• {{cite journal|last1=Cutler|first1=David F.|last2=Gregory|first2=Mary|title=Current anatomical research in Liliaceae, Amaryllidaceae and Iridaceae|journal=Telopea|date=13 Oct 1983|volume=2|issue=4|pages=425–452|url=https://www.rbgsyd.nsw.gov.au/science/Scientific_publications/telopea/view_contents/b-view_contents?sq_content_src=%2BdXJsPWh0dHAlM0ElMkYlMkZwbGFudG5ldC5yYmdzeWQubnN3Lmdvdi5hdSUyRmVtdXdlYm5zd2xpdmUlMkZvYmplY3RzJTJGY29tbW9uJTJGd2VibWVkaWEucGhwJTNGaXJuJTNENzU1OTYlMjZyZWZ0YWJsZSUzRGViaWJsaW9ncmFwaHkmYWxsPTE%3D|doi=10.7751/telopea19834408 |accessdate=26 January 2015}}
• {{cite journal|last1=Fay|first1=Michael F.|last2=Chase|first2=Mark W.|authorlink1=Michael F Fay|authorlink2=Mark W Chase|title=Resurrection of Themidaceae for the Brodiaea alliance, and Recircumscription of Alliaceae, Amaryllidaceae and Agapanthoideae|journal=Taxon|date=August 1996|volume=45|issue=3|pages=441–451|ref={{harvid|Fay|Chase|1996}}|doi=10.2307/1224136|jstor=1224136}}
• {{cite journal|last1=García|first1=Nicolás|last2=Meerow|first2=Alan W.|last3=Soltis|first3=Douglas E.|last4=Soltis|first4=Pamela S.|authorlink2=Alan Meerow|authorlink4 = Pamela S. Soltis|title=Testing Deep Reticulate Evolution in Amaryllidaceae Tribe Hippeastreae (Asparagales) with ITS and Chloroplast Sequence Data|journal=Systematic Botany|date=1 March 2014|volume=39|issue=1|pages=75–89|doi=10.1600/036364414X678099|ref={{harvid|Garcia et al|2014}}}}
• {{cite journal | last1 = Meerow | first1 = A.W. | last2 = Fay | first2 = M.F. | last3 = Guy | first3 = C.L. | last4 = Li | first4 = Q.-B. | last5 = Zaman | first5 = F.Q. | last6 = Chase | first6 = M.W. | authorlink1=Alan Meerow|authorlink2=Michael F Fay|authorlink6=Mark W Chase|title = Systematics of Amaryllidaceae based on cladistic analysis of plastid rbcL and trnL-F sequence data | journal = Am. J. Bot. |year = 1999 | volume = 86 | issue = 9 | pages = 1325–1345 | doi = 10.2307/2656780|ref={{harvid|Meerow et al.|1999}} | pmid=10487820 | jstor=2656780}}
• {{cite journal | last1 = Meerow | first1 = A.W. | last2 = Guy | first2 = C.L. | last3 = Li | first3 = Q.-B. | last4 = Yang | first4 = S.-L. | authorlink1 = Alan Meerow | title = Phylogeny of the American Amaryllidaceae Based on nrDNA ITS Sequences | journal = Systematic Botany | year = 2000 | volume = 25 | issue = 4 | pages = 708–726 | doi = 10.2307/2666729 | url = http://www.bulbsociety.org/meerow/Meerow%20et%20al-American.pdf | accessdate = 25 January 2015 | ref = {{harvid|Meerow et al.|2000b}} | jstor = 2666729 | archive-url = https://web.archive.org/web/20150923195144/http://www.bulbsociety.org/meerow/Meerow%20et%20al-American.pdf | archive-date = 2015-09-23 | dead-url = yes | df = }}
• {{cite journal|last1=Meerow|first1=Alan W.|last2=Snijman|first2=Deirdre A.|authorlink1=Alan Meerow|title=Phylogeny of Amaryllidaceae Tribe Amaryllideae Based on nrDNA ITS Sequences and Morphology|journal=American Journal of Botany|date=December 2001|volume=88|issue=12|pages=2321–2330|doi=10.2307/3558392|url=https://naldc-legacy.nal.usda.gov/naldc/download.xhtml?id=3004&content=PDF|jstor=3558392}}
• {{cite journal|last1=Meerow|first1=Alan W.|last2=Guy|first2=Charles L.|last3=Li|first3=Qin-Bao|last4=Clayton|first4=Jason R.|authorlink1=Alan Meerow|title=Phylogeny of the Tribe Hymenocallideae (Amaryllidaceae) Based on Morphology and Molecular Characters|journal=Annals of the Missouri Botanical Garden|date=2002|volume=89|issue=3|pages=400–413|doi=10.2307/3298600|jstor=3298600|url=https://www.academia.edu/6359033}}
• {{cite journal|last1=Meerow|first1=Alan W.|last2=Clayton|first2=Jason R.|authorlink1=Alan Meerow|title=Generic relationships among the baccate-fruited Amaryllidaceae (tribe Haemantheae) inferred from plastid and nuclear non-coding DNA sequences|journal=Plant Systematics and Evolution|date=1 February 2004|volume=244|issue=3–4|pages=141–155|doi=10.1007/s00606-003-0085-z}}
• {{cite journal|last1=Meerow|first1=Alan W.|last2=van der Werff|first2=Henk|authorlink1=Alan Meerow|title=Pucara (Amaryllidaceae) Reduced to Synonymy with Stenomesson on the Basis of Nuclear and Plastid DNA Spacer Sequences, and a New Related Species of Stenomesson|url=https://www.researchgate.net/publication/266271150|journal=Systematic Botany|date=1 July 2004|volume=29|issue=3|pages=511–517|doi=10.1600/0363644041744400|ref={{harvid|Meerow et al|2004}}}}
• {{cite journal|last1=Meerow|first1=AW|last2=Francisco-Ortega|first2=J|last3=Schnell|first3=RJ|authorlink1=Alan Meerow|title=Phylogenetic relationships and biogeography within the Eurasian clade of Amaryllidaceae based on plastid ndhF and nrDNA ITS sequences: lineage sorting in a reticulate area?|journal=Systematic Botany|date=2006|volume=31|pages=42–60|doi=10.1600/036364406775971787|jstor=25064128|issue=1|ref={{harvid|Meerow et al|2006a}}}} Full text
• {{cite journal|last1=Meerow|first1=Alan W.|last2=Snijman|first2=Deirdre A.|authorlink1=Alan Meerow|title=The never-ending story: multigene approaches to the phylogeny of Amaryllidaceae|journal=Aliso|date=2006|volume=22|pages=355–366|url=http://naldc-legacy.nal.usda.gov/naldc/download.xhtml?id=2425&content=PDF|accessdate=25 January 2015|ref=harv|df=|doi=10.5642/aliso.20062201.29}}
• {{cite journal|last1=Meerow|first1=Alan W.|last2=Reveal|first2=James L.|last3=Snijman|first3=Deirdre A.|last4=Dutilh|first4=Julie H.|authorlink1=Alan Meerow|title=(1793) Proposal to conserve the name Amaryllidaceae against Alliaceae, a "superconservation" proposal|journal=Taxon|date=November 2007|volume=56|issue=4|pages=1299–1300|url=http://www.ingentaconnect.com/content/iapt/tax/2007/00000056/00000004/art00029|accessdate=28 January 2015|ref={{harvid|Meerow et al.|2007}}|doi=10.2307/25065925|jstor=25065925}}
• {{cite journal|last1=Müller-Doblies|first1=Dietrich|last2=Müller-Doblies|first2=Ute|authorlink1=Dietrich Müller-Doblies|authorlink2=Ute Müller-Doblies|title=Studies on tribal systematics of Amaryllidoideae: 1. The systematic position of Lapiedra Lag|journal=Lagascalia|date=1978|volume=8|issue=1|pages=13–23|url=https://www.researchgate.net/publication/28292204}}
• {{cite journal|last1=Müller-Doblies|first1=U.|last2=Müller-Doblies|first2=D.|authorlink2=Dietrich Müller-Doblies|authorlink1=Ute Müller-Doblies|title=Tribes and subtribes and some species combinations in Amaryllidaceae J St Hil R Dahlgren & al. 1985|journal=Feddes Repertorium|date=1996|volume=107|issue=5–6|pages=S.c.1–S.c.9|ref=harv}}
• {{Cite book|last1=Thorne|first1=Robert F.|authorlink=Robert F Thorne|title=A Phylogenetic Classification of the Angiospermae|journal=Evolutionary Biology|date=1976|volume=9|pages=35–106|doi=10.1007/978-1-4615-6950-3_2|ref=harv|isbn=978-1-4615-6952-7}}
• {{cite journal |last=Thorne |first=R. F. |authorlink=Robert F Thorne|title=Classification and geography of the flowering plants |journal=The Botanical Review |year=1992 |volume=58 |issue=3 |pages=225–348 |doi=10.1007/BF02858611 |ref=harv}}
• {{cite journal|last1=Weber|first1=Odile|last2=Wilkin|first2=Paul|title=588. Stenomesson pearcei|journal=Curtis's Botanical Magazine|date=May 2007|volume=24|issue=2|pages=114–120|doi=10.1111/j.1467-8748.2007.00572.x|ref=harv}}
• {{cite book|last1=Strydom|first1=Aéle|title=Phylogenetic relationships in the family Amaryllidaceae|url=http://etd.uovs.ac.za/ETD-db//theses/available/etd-09122006-103922/unrestricted/StrydomA.pdf |date=2005|publisher=Department of Plant Sciences, University of the Free State|format=PhD thesis}}

APG system

• {{cite journal|author = Angiosperm Phylogeny Group|authorlink=Angiosperm Phylogeny Group|year = 1998|title = An ordinal classification for the families of flowering plants|journal = Annals of the Missouri Botanical Garden|volume = 85|pages = 531–553|doi = 10.2307/2992015|jstor = 2992015|issue = 4|ref={{harvid|APG|1998}}}}
• {{cite journal|author = Angiosperm Phylogeny Group|authorlink=Angiosperm Phylogeny Group|title=An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG II|journal=Botanical Journal of the Linnean Society|volume = 141|issue=4|pages = 399–436|date=April 2003|doi=10.1046/j.1095-8339.2003.t01-1-00158.x|ref={{harvid|APG|2003}}}}
• {{Citation |last=Angiosperm Phylogeny Group |authorlink=Angiosperm Phylogeny Group|year=2009 |title=An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III |journal=Botanical Journal of the Linnean Society |volume=161 |issue=2 |pages=105–121 |url=http://www3.interscience.wiley.com/journal/122630309/abstract |archive-url=https://wayback.archive-it.org/all/20170525104318/http://onlinelibrary.wiley.com/doi/10.1111/j.1095-8339.2009.00996.x/abstract |dead-url=yes |archive-date=25 May 2017 |accessdate= 7 January 2015 |doi=10.1111/j.1095-8339.2009.00996.x|ref={{harvid|APG|2009}}}}
• {{cite journal |last = APG IV |authorlink=Angiosperm Phylogeny Group|title=An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG IV |year=2016 |journal=Botanical Journal of the Linnean Society |volume=181 |issue=1 |pages=1–20 |doi=10.1111/boj.12385 |ref=harv}}
• {{Citation | last = Chase | first = M.W. |last2 = Reveal | first2 = J.L. | last3 = Fay | first3 = M.F. | authorlink1=Mark W Chase|authorlink2=James Reveal |authorlink3=Michael F. Fay|year = 2009 | title = A subfamilial classification for the expanded asparagalean families Amaryllidaceae, Asparagaceae and Xanthorrhoeaceae | journal = Botanical Journal of the Linnean Society | volume = 161 | issue = 2 | pages = 132–136 | doi = 10.1111/j.1095-8339.2009.00999.x|ref={{harvid|Chase et al.|2009}} }}

Pharmacology

• {{cite journal | last1 = Ronsted | first1 = N | last2 = Symonds | first2 = MR | last3 = Birkholm | first3 = T | last4 = Christensen | first4 = SB | last5 = Meerow | first5 = AW | display-authors = etal | year = 2012 | title = Can phylogeny predict chemical diversity and potential medicinal activity of plants? A case study of Amaryllidaceae | url = | journal = BMC Evol Biol | volume = 12 | issue = 1| page = 182 | doi = 10.1186/1471-2148-12-182 | pmid = 22978363 | pmc = 3499480 }}
• {{cite journal|last1=Takos|first1=Adam|last2=Rook|first2=Fred|title=Towards a Molecular Understanding of the Biosynthesis of Amaryllidaceae Alkaloids in Support of Their Expanding Medical Use|journal=International Journal of Molecular Sciences|date=31 May 2013|volume=14|issue=6|pages=11713–11741|doi=10.3390/ijms140611713|url=http://www.mdpi.com/1422-0067/14/6/11713/htm|accessdate=25 December 2014|ref=harv|pmid=23727937|pmc=3709753}}

Websites

• {{cite web|last1=Vigneron|first1=Pascal|title=Amaryllidaceae|url=http://www.amaryllidaceae.org//index.htm|website=Amaryllidaceae.org|accessdate=23 October 2014|language=French}}
• {{cite web|last1=Meerow|first1=A|title=Neotropical Amaryllidaceae|url=http://www.kew.org/science/tropamerica/neotropikey/families/Amaryllidaceae.htm|website=Milliken, W., Klitgård, B. & Baracat, A. Neotropikey - Interactive key and information resources for flowering plants of the Neotropics|accessdate=8 January 2015|date=2009}}
• {{cite web|last1=Dutilh|first1= J.H.A.| title=Neotropical Alliaceae|url=http://www.kew.org/science/tropamerica/neotropikey/families/Alliaceae.htm|website=Milliken, W., Klitgård, B. & Baracat, A. Neotropikey - Interactive key and information resources for flowering plants of the Neotropics|accessdate=8 January 2015|date=2009}}
• {{cite web|title=Linnaeus Sexual System|url=http://cronklab.wikidot.com/linnaeus-sexual-system|website=CronkLab|publisher=Biodiversity Research Centre, University of British Columbia|accessdate=26 January 2015|ref={{harvid|Linnaeus Sexual System|2015}}}}
• {{cite web|last=ICN|authorlink=International Code of Nomenclature for algae, fungi, and plants|title=International Code of Nomenclature for algae, fungi, and plants|url=http://www.iapt-taxon.org/nomen/main.php?page=title|publisher=International Association for Plant Taxonomy|accessdate=2 February 2014|location=Bratislava|year=2011|ref=harv}}
• {{cite web|title=Pacific Bulb Society|url=http://www.pacificbulbsociety.org/|publisher=Pacific Bulb Society|accessdate=9 February 2016|date=2012|ref={{harvid|PBS|2012}}}}

Databases

• {{cite web|title=Amaryllidaceae|url=http://www.theplantlist.org/1.1/browse/A/Amaryllidaceae/|website=The Plant List (2013). Version 1.1.|accessdate=4 January 2015|date=2013|ref={{harvid|The Plant List|2013}}}}
• {{cite web |title=Quick Search|website=World Checklist of Selected Plant Families |publisher=Royal Botanic Gardens, Kew |url=http://apps.kew.org/wcsp/home.do|accessdate=8 January 2015 }}
• {{citation |last=Stevens |first=P.F. |authorlink=Peter F Stevens|date=2016|origyear= 2001 |title=Angiosperm Phylogeny Website |url=http://www.mobot.org/mobot/research/apweb/|publisher=Missouri Botanical Gardens|accessdate=7 February 2016|ref={{harvid|Stevens|2016}}}}
• {{cite web|last1=Wilkin|first1=Paul|title=Amaryllidaceae J.St.-Hil|url=http://e-monocot.org/taxon/urn:kew.org:wcs:family:81|website=eMonocot|accessdate=27 January 2015|date=2012|ref=harv}}
• {{cite web|title=Amaryllidaceae: A taxonomic tool for the Amaryllidaceae of the world|url=http://amaryllidaceae.e-monocot.org/|publisher=eMonocot|ref={{harvid|eMonocot|2016}}}}
• {{IPNI|id= 30000959-2 }}

External links

• Amaryllidaceae J.St.-Hil. at the Amaryllidaceae project, e-monocot.org
• Amaryllidaceae in BoDD – Botanical Dermatology Database

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