| 词条 | Phyllanthus |
| 释义 |
|image = Phyllanthus mirabilis5 ies.jpg |image_caption = Phyllanthus mirabilis |display_parents = 3 |taxon = Phyllanthus |authority = L. |synonyms = {{hidden begin|toggle=left|title=List}}
|synonyms_ref = [1] |diversity = About 800 species |diversity_link = List of Phyllanthus species |subdivision_ranks = Species }} Phyllanthus is the largest genus in the flowering plant family Phyllanthaceae. Estimates of the number of species in this genus vary widely, from 750[2] to 1200.[3] Phyllanthus has a remarkable diversity of growth forms including annual and perennial herbs, shrubs, climbers, floating aquatics, and pachycaulous succulents. Some have flattened leaflike stems called cladodes. It has a wide variety of floral morphologies and chromosome numbers and has one of the widest range of pollen types of any seed plant genus. Despite their variety, almost all Phyllanthus species express a specific type of growth called "phyllanthoid branching" in which the vertical stems bear deciduous, floriferous (flower-bearing), plagiotropic (horizontal or oblique) stems. The leaves on the main (vertical) axes are reduced to scales called "cataphylls", while leaves on the other axes develop normally.[4] Phyllanthus is distributed in all tropical and subtropical regions on Earth. SpeciesThe circumscription of this genus has been a cause of much confusion and disagreement. Molecular phylogenetic studies have shown that Phyllanthus is paraphyletic over Reverchonia, Glochidion, Sauropus, and Breynia. A recent revision of the family Phyllanthaceae has subsumed all four of these genera into Phyllanthus.[5] This enlarged version of Phyllanthus might eventually be divided into smaller genera,[6][7] including 32 Chinese (and northern Indochinese) species.[8]
For full list, see List of Phyllanthus species. ==Fossil record== Two fossil seeds of a Phyllanthus species have been extracted from borehole samples of the Middle Miocene fresh water deposits in Nowy Sacz Basin, West Carpathians, Poland. The seeds are similar to seeds of the fossil species †Phyllanthus triquetra and †Phyllanthus compassica from the Oligocene and Miocene of West Siberia. Phyllanthus fossils are known from several Miocene and Pliocene sites in Poland.[12] Pollination biologyPhyllanthus are of note in the fields of pollination biology and coevolution because some but not all species in the genus have a specialized mutualism with moths in the genus Epicephala (leafflower moths), in which the moths actively pollinate the flowers. While ensuring that the tree may produce viable seeds, the moths also lay eggs in the flowers' ovaries where their larvae consume a subset of the developing seeds as nourishment.[13][14] Other species of Epicephala are pollinators of certain species of plants in the genera Glochidion[15][16] and Breynia,[17][18] both of which are phylogenetically nested within Phyllanthus.[19]Research and traditional medicineParticularly for its content of tannins, P. emblica fruit has a history of use in traditional medicine and is under study for its potential biological properties.[20] Leaves, roots, stem, bark and berries of this genus contain lignans and other phytochemicals.[21][22][23] Phyllanthus species have been assessed for use in people with chronic hepatitis B virus infection, but the low quality of clinical trials has prevented any conclusion about efficacy as of 2013.[24][25]References1. ^{{cite web|url=http://apps.kew.org/wcsp/synonomy.do?name_id=153270|title=World Checklist of Selected Plant Families}} {{Commons category}}2. ^David J. Mabberley. 2008. Mabberley's Plant-Book. third edition (2008). Cambridge University Press.{{Page needed|date=September 2010}} 3. ^{{Cite journal|vauthors=Kathriarachchi H, Hoffmann P, Samuel R, Wurdack KJ, Chase MW |title=Molecular phylogenetics of Phyllanthaceae inferred from five genes (plastid atpB, matK, 3'ndhF, rbcL, and nuclear PHYC) |journal=Molecular Phylogenetics and Evolution |volume=36 |issue=1 |pages=112–34 |date=July 2005 |pmid=15904861 |doi=10.1016/j.ympev.2004.12.002}} 4. ^{{Cite journal|first=Grady L. |last=Webster |year=1994 |title=Classification of the Euphorbiaceae |journal=Annals of the Missouri Botanical Garden |volume=81 |issue=1 |pages=3–32 |doi=10.2307/2399908 |jstor=2399908}} 5. ^{{cite journal | last1 = Hoffmann | first1 = Petra | last2 = Kathriarachchi | first2 = Hashendra S. | last3 = Wurdack | first3 = Kenneth J. | year = 2006 | title = A Phylogenetic Classification of Phyllanthaceae | url = | journal = Kew Bulletin | volume = 61 | issue = 1| pages = 37–53 }} 6. ^{{cite journal | doi = 10.3732/ajb.93.4.637 | last1 = Kathriarachchi | first1 = Hashendra S. | last2 = Samuel | first2 = Rosabelle | last3 = Hoffmann | first3 = Petra | last4 = Mlinarec | first4 = Jelena | last5 = Wurdack | first5 = Kenneth J. | last6 = Ralimanana | first6 = Hélène | last7 = Stuessy | first7 = Tod F. | last8 = Chase | first8 = Mark W. | year = 2006 | title = Phylogenetics of tribe Phyllantheae (Phyllanthaceae) based on nrITS and plastid matK DNA sequence data | url = | journal = American Journal of Botany | volume = 93 | issue = 4| pages = 637–655 | pmid = 21646224 }} 7. ^Kanchana Pruesapan, Ian R.H. Telford, Jeremy J. Bruhl, Stefano G.A. Draisma, and Peter C. Van Welzen. 2008. "Delimitation of Sauropus (Phyllanthaceae) Based on Plastid matK and Nuclear Ribosomal ITS DNA Sequence Data." Annals of Botany 102(6):1007-1018 8. ^{{cite web|url=http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=125179|title=Flora of China|publisher=eFlora|accessdate=18 April 2017}} 9. ^[https://www.jstor.org/stable/26296180?seq=1#page_scan_tab_contents "Phyllanthus phialanthoides (Phyllanthaceae), a new species from northeastern Cuba"] (JSTOR) 10. ^1 Luo, S.X., H.-J. Esser, D. Zhang, and S. S. Renner. 2011. Nuclear ITS sequences help disentangle Phyllanthus reticulatus (Phyllanthaceae), an Asian species not occurring in Africa, but introduced to Jamaica. Systematic Botany 36(1): 99-104. 11. ^{{cite journal | doi = 10.1186/1746-4269-2-22 | last1 = Bussmann | first1 = R. W. | last2 = Gilbreath | year = 2006 | first2 = GG | last3 = Solio | first3 = J | last4 = Lutura | first4 = M | last5 = Lutuluo | first5 = R | last6 = Kunguru | first6 = K | last7 = Wood | first7 = N | last8 = Mathenge | first8 = SG | title = Plant use of the Maasai of Sekenani Valley, Maasai Mara, Kenya | journal = J Ethnobiol Ethnomed | volume = 2 | issue = | page = 22 | pmid = 16674830 | pmc = 1475560 }} 12. ^Macroscopic plant remains from the freshwater Miocene of the Nowy Sącz Basin (West Carpathians, Poland) by Łańcucka-Środoniowa M, Acta Palaeobotanica 1979 20 (1): 3-117. 13. ^Kawakita, A.; Kato, M. 2004. "Evolution of obligate pollination mutualism in New Caledonian Phyllanthus (Euphorbiaceae)." American Journal of Botany 91: 410–415. 14. ^Kawakita, A.; Kato, M. 2009. "Repeated independent evolution of obligate pollination mutualism in the Phyllantheae-Epicephala association." Proceedings of the Royal Society B. 276: 417–426. 15. ^Hembry, D. H.; Okamoto, T.; Gillespie, R. G. (2012) Repeated colonization of remote islands by specialized mutualists. Biology Letters. 8: 258–261. 16. ^Luo, S.-X.; Yao, G.; Wang, Z.; Zhang, D.; Hembry, D. H. (2017) "A novel, enigmatic basal leafflower moth lineage pollinating a derived leafflower host illustrates the dynamics of host shifts, partner replacement, and apparent co-adaptation in intimate mutualisms." The American Naturalist. 189: 422–435 17. ^Kawakita, A.; Kato, M. 2004. Obligate pollination mutualism in Breynia (Phyllanthaceae): further documentation of pollination mutualism involving Epicephala moths (Gracillariidae). American Journal of Botany. 91: 1319–1325. 18. ^Zhang, J.; Wang, S.; Li, H.; Hu, B.; Yang, X.; Wang, Z. 2012. "Diffuse coevolution between two Epicephala species (Gracillariidae) and two Breynia species (Phyllanthaceae). PLOS ONE. 7: e41657. 19. ^Kathriarachchi, H.; Samuel, R.; Hoffmann, P.; Mlinarec, J.; Wurdack, K. J.; Ralimanana, H.; Stuessy, T. F.; Chase, M. W. 2006. "Phylogenetics of tribe Phyllantheae (Phyllanthaceae: Euphorbiaceae sensu lato) based on nrITS and plastid matK DNA sequence data." American Journal of Botany. 93: 637–655. 20. ^{{cite journal|pmid=24369850|year=2014|author1=Yang|first1=B|title=Composition and biological activities of hydrolyzable tannins of fruits of Phyllanthus emblica|journal=Journal of Agricultural and Food Chemistry|volume=62|issue=3|pages=529–41|last2=Liu|first2=P|doi=10.1021/jf404703k}} 21. ^{{Cite journal|vauthors=Murugaiyah V, Chan KL |title=Determination of four lignans in Phyllanthus niruri L. by a simple high-performance liquid chromatography method with fluorescence detection |journal=Journal of Chromatography A |volume=1154 |issue=1–2 |pages=198–204 |date=June 2007 |pmid=17418855 |doi=10.1016/j.chroma.2007.03.079}} 22. ^{{Cite journal |vauthors=Srivastava V, Singh M, Malasoni R, etal |title=Separation and quantification of lignans in Phyllanthus species by a simple chiral densitometric method |journal=Journal of Separation Science |volume=31 |issue=1 |pages=47–55 |date=January 2008 |pmid=18064620 |doi=10.1002/jssc.200700282}} 23. ^{{Cite journal|vauthors=Bagalkotkar G, Sagineedu SR, Saad MS, Stanslas J |title=Phytochemicals from Phyllanthus niruri Linn. and their pharmacological properties: a review |journal=The Journal of Pharmacy and Pharmacology |volume=58 |issue=12 |pages=1559–70 |date=December 2006 |pmid=17331318 |doi=10.1211/jpp.58.12.0001}} 24. ^{{cite journal|pmid=11555193|year=2001|author1=Liu|first1=J|title=Genus Phyllanthus for chronic hepatitis B virus infection: A systematic review|journal=Journal of Viral Hepatitis|volume=8|issue=5|pages=358–66|last2=Lin|first2=H|last3=McIntosh|first3=H|doi=10.1046/j.1365-2893.2001.00307.x}} 25. ^{{cite journal|pmid=23633363|year=2013|author1=Xia|first1=Y|title=Phyllanthus species versus antiviral drugs for chronic hepatitis B virus infection|journal=The Cochrane Database of Systematic Reviews|issue=4|pages=CD009004|last2=Luo|first2=H|last3=Liu|first3=J. P.|last4=Gluud|first4=C|doi=10.1002/14651858.CD009004.pub2}} External links
2 : Phyllanthus|Malpighiales genera |
| 随便看 |
|
开放百科全书收录14589846条英语、德语、日语等多语种百科知识，基本涵盖了大多数领域的百科知识，是一部内容自由、开放的电子版国际百科全书。