词条 | Pomacentridae |
释义 |
| name = Clownfish and damselfish | image = cocoa damselfish.jpg | image_caption = Cocoa damselfish, Stegastes variabilis | taxon = Pomacentridae | authority = Bonaparte, 1832 | subdivision_ranks = Genera | subdivision = See text. }}{{Commons category|Pomacentridae}}Pomacentridae is a family of ray-finned fish, comprising the damselfishes and clownfishes. This family were formerly placed in the order Perciformes but are now regarded as being incertae sedis in the subseries Ovalentaria in the clade Percomorpha.[1] They are primarily marine, while a few species inhabit freshwater and brackish environments (e.g., Neopomacentrus aquadulcis, N. taeniurus, Pomacentrus taeniometopon, Stegastes otophorus).[2] They are noted for their hardy constitutions and territoriality. Many are brightly colored, so they are popular in aquaria. Around 385 species are classified in this family, in about 29 genera. Of these, members of two genera, Amphiprion and Premnas (subfamily Amphiprioninae), are commonly called clownfish or anemonefish, while members of other genera (e.g., Pomacentrus) are commonly called damselfish.[3] The members of this family are classified in four subfamilies: Amphiprioninae, Chrominae, Lepidozyginae, and Pomacentrinae.[4] EtymologyThe name of the family is derived from the Greek words; poma roughly translates to the English "cover", referring to the fishes' opercula, and kentron is Greek for sting. The name refers to the serrations found along the margins of the opercular bones in many members of this family.[3] Distribution and habitatPomacentrids are found primarily in tropical seas, with a few species occurring in temperate waters (e.g., Hypsypops rubicundus). Most species are found on or near coral reefs in the Indo-West Pacific (from East Africa to Polynesia). The area from the Philippines to Australia hosts the greatest concentration of species.[5] The remaining species are found in the Atlantic or eastern Pacific.{{citation needed|date=January 2015}} Some species are native to freshwater or brackish estuarine environments.[2][6] Most members of the family live in shallow water, from {{convert|2|to|15|m|abbr=on}} in depth, although some species (e.g., Chromis abyssus) are found below {{convert|100|m|ft|abbr=on}}.[7] Most species are specialists, living in specific parts of the reef, such as sandy lagoons, steep reef slopes, or areas exposed to strong wave action. In general, the coral is used as shelter, and many species can only survive in its presence.[6] The bottom-dwelling species are territorial, occupying and defending a portion of the reef, often centered on shelter. By keeping away other species of fish, some pomacentrids encourage the growth of thick mats of algae within their territories, leading to the common name farmerfish.[6] CharacteristicsPomacentrids have an orbiculate to elongated body shape, which is often laterally compressed. They have interrupted or incomplete lateral lines and they usually have a single nostril on each side (some species of Chromis and Dascyllus have two on each side).[5] They have small- to medium-sized ctenoid scales. They have one or two rows of teeth, which may be conical or spatulate. They display a wide range of colors, predominantly bright shades of yellow, red, orange, and blue, although some are a relatively drab brown, black, or grey. The young are often a different, brighter color than adults. Pomacentrids are omnivorous or herbivorous, feeding on algae, plankton, and small bottom-dwelling crustaceans, depending on their precise habitats. Only a small number of genera, such as Cheiloprion, eat the coral where they live.[6] They also engage in symbiotic relationship with cleaner gobies of genus Elacatinus, allowing the gobies to feed on ectoparasites on their bodies.[8] LifecycleBefore breeding, the males clear an area of algae and invertebrates to create a nest. They engage in ritualised courtship displays, which may consist of rapid bursts of motion, chasing or nipping females, stationary hovering, or wide extension of their fins. After being attracted to the site, the female lays a string of sticky eggs that attach to the substrate. The male swims behind the female as she lays the eggs, and fertilises them externally. Varying by species, brood sizes range from 50 to 1000 eggs.[6] The male guards the nest for the two to seven days needed for the eggs to hatch. The transparent larvae are {{convert|2|to|4|mm|abbr=on}} long. They go through a pelagic stage, which depending on the species, can last as little as a week or more than a month.[9] When they arrive at a suitable environment, the young settle and adopt their juvenile colors.[6] In captivity, pomacentrids live up to 18 years, but they probably do not live longer than 10 to 12 years in the wild.[6] GeneraAbout 28 genera in four subfamilies are recognized:[10] {{col-begin|width=auto}}{{col-break}}
TimelineImageSize = width:1000px height:auto barincrement:15px PlotArea = left:10px bottom:50px top:10px right:10px Period = from:-65.5 till:10 TimeAxis = orientation:horizontal ScaleMajor = unit:year increment:5 start:-65.5 ScaleMinor = unit:year increment:1 start:-65.5 TimeAxis = orientation:hor AlignBars = justify Colors = #legends id:CAR value:claret id:ANK value:rgb(0.4,0.3,0.196) id:HER value:teal id:HAD value:green id:OMN value:blue id:black value:black id:white value:white id:cenozoic value:rgb(0.54,0.54,0.258) id:paleogene value:rgb(0.99,0.6,0.32) id:paleocene value:rgb(0.99,0.65,0.37) id:eocene value:rgb(0.99,0.71,0.42) id:oligocene value:rgb(0.99,0.75,0.48) id:neogene value:rgb(0.999999,0.9,0.1) id:miocene value:rgb(0.999999,0.999999,0) id:pliocene value:rgb(0.97,0.98,0.68) id:quaternary value:rgb(0.98,0.98,0.5) id:pleistocene value:rgb(0.999999,0.95,0.68) id:holocene value:rgb(0.999,0.95,0.88) BarData= bar:eratop bar:space bar:periodtop bar:space bar:NAM1 bar:space bar:period bar:space bar:era PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 shift:(7,-4) bar:periodtop from: -65.5 till: -55.8 color:paleocene text:Paleocene from: -55.8 till: -33.9 color:eocene text:Eocene from: -33.9 till: -23.03 color:oligocene text:Oligocene from: -23.03 till: -5.332 color:miocene text:Miocene from: -5.332 till: -2.588 color:pliocene text:Plio. from: -2.588 till: -0.0117 color:pleistocene text:Pleist. from: -0.0117 till: 0 color:holocene text:H. bar:eratop from: -65.5 till: -23.03 color:paleogene text:Paleogene from: -23.03 till: -2.588 color:neogene text:Neogene from: -2.588 till: 0 color:quaternary text:Q. PlotData= PlotData= bar:period from: -65.5 till: -55.8 color:paleocene text:Paleocene from: -55.8 till: -33.9 color:eocene text:Eocene from: -33.9 till: -23.03 color:oligocene text:Oligocene from: -23.03 till: -5.332 color:miocene text:Miocene from: -5.332 till: -2.588 color:pliocene text:Plio. from: -2.588 till: -0.0117 color:pleistocene text:Pleist. from: -0.0117 till: 0 color:holocene text:H. bar:era from: -65.5 till: -23.03 color:paleogene text:Paleogene from: -23.03 till: -2.588 color:neogene text:Neogene from: -2.588 till: 0 color:quaternary text:Q. References1. ^{{cite book |title=Fishes of the World |edition=5th |author1=J. S. Nelson |author2=T. C. Grande |author3=M. V. H. Wilson |year=2016 |pages=752 |publisher=Wiley |isbn= 978-1-118-34233-6 |url=https://sites.google.com/site/fotw5th/ }} 2. ^1 {{cite journal |author1=Jenkins, A.P. |author2=G.R. Allen |lastauthoramp=yes | year=2002 | title=Neopomacentrus aquadulcis, a new species of damselfish (Pomacentridae) from eastern Papua New Guinea | journal=Records of the Western Australian Museum | volume=20 | pages=379–382}} 3. ^1 {{FishBase family | family = Pomacentridae | year = 2007 | month = July}} 4. ^{{cite book | author=Allen, G.R. | year=1975 | title=Damselfishes of the South Seas | publisher=T.F.H. Publications | location=Neptune City, NJ | isbn=978-0-87666-034-8}} 5. ^1 {{cite book | last=Nelson |first=J.S. | year=2006 | title=Fishes of the World | publisher=John Wiley and Sons | location=Hoboken | isbn=978-0-471-25031-9}} 6. ^1 2 3 4 5 6 {{cite book |editor=Paxton, J.R. |editor2=Eschmeyer, W.N.|author= Allen, Gerald R.|year=1998|title=Encyclopedia of Fishes|publisher= Academic Press|location=San Diego|pages= 205–208|isbn= 978-0-12-547665-2}} 7. ^{{cite journal | author=Pyle, R.L., J.L. Earle & B.D. Greene | year=2008 | title=Five new species of the damselfish genus Chromis (Perciformes: Labroidei: Pomacentridae) from deep coral reefs in the tropical western Pacific | journal=Zootaxa | volume=1671 | pages=3–31 | url=http://plazi.org:8080/dspace/bitstream/10199/15417/6/zt01671p031.pdf | deadurl=yes | archiveurl=https://web.archive.org/web/20110721153207/http://plazi.org:8080/dspace/bitstream/10199/15417/6/zt01671p031.pdf | archivedate=2011-07-21 | df= }} 8. ^{{Cite journal|author1=Ivan Sazima |author2=Cristina Sazima |author3=Ronaldo B. Francini-Filho |author4=Rodrigo L. Moura |date=September 2000| title = Daily cleaning activity and diversity of clients of the barber goby, Elacatinus figaro, on rocky reefs in southeastern Brazil| journal = Environmental Biology of Fishes | volume =59 | issue =1| pages =69–77 | doi=10.1023/a:1007655819374}} 9. ^{{cite journal | last1=Thresher |first1=R.E. |first2=P.L. |last2=Colin |first3=L.J. |last3=Bell | year=1989 | title=Planktonic duration, distribution and population structure of western and central Pacific damselfishes (Pomacentridae) | journal=Copeia | volume=1989 | issue=2 | pages=420–434 | jstor=1445439 | doi=10.2307/1445439}} 10. ^{{Cite book| edition = 4| publisher = John Wiley & Sons| isbn = 978-0-471-25031-9| last = Nelson| first = J. S.| title = Fishes of the World| authorlink = Joseph S. Nelson| location = Hoboken, NJ| date = 2006| title-link = Fishes of the World}} External links{{Wikispecies|Pomacentridae}}
5 : Pomacentridae|Labroidei|Ovalentaria|Marine fish families|Taxa named by Charles Lucien Bonaparte |
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