词条 | Saxifraga | ||||||
释义 |
|image = Saxifraga cochlearis1.jpg |image_caption = Saxifraga cochlearis |taxon = Saxifraga |authority = L. |subdivision_ranks = Sections |subdivision =
|synonyms = Boecherarctica Á.Löve Cascadia A.M.Johnson Micranthes Haw. Zahlbrucknera Rchb. |synonyms_ref = [1] }}Saxifraga is the largest genus in the family Saxifragaceae, containing about 440 species of holarctic perennial plants, known as saxifrages[2] or rockfoils.[3] The Latin word saxifraga means literally "stone-breaker", from Latin {{lang|la|saxum}} ("rock" or "stone") + {{lang|la|frangere}} ("to break"). It is usually thought to indicate a medicinal use for treatment of urinary calculi (known as kidney stones), rather than breaking rocks apart.[2][4] The genera Saxifragopsis (strawberry saxifrage), and Saxifragella are sometimes included in Saxifraga.[1] In recent DNA based phylogenetic analyses of the Saxifragaceae, the former sections Micranthes and Merkianae are shown to be more closely related to the Boykinia and Heuchera clades,[5] and the most recent floras separate these groups as the genus Micranthes.[6][7] DescriptionMost saxifrages are smallish plants whose leaves grow close to the ground, often in a rosette. The leaves typically have a more or less incised margin; they may be succulent, needle-like and/or hairy, reducing evaporation.[10] The inflorescence or single flower clusters rise above the main plant body on naked stalks. The small actinomorphic hermaphrodite flowers have five petals and sepals and are usually white, but red to yellow in some species.{{citation needed|date=October 2011}} As in other primitive eudicots, some of the 5 or 10 stamens may appear petal-like.{{citation needed|date=October 2011}} and it lives tundra ecosystem. Ecology{{Refimprove section|date = November 2018}}Saxifrages are typical inhabitants of Arctic–alpine ecosystems, and are hardly ever found outside the temperate parts of the Northern Hemisphere; most members of this genus are found in subarctic climates. A good number of species grow in glacial habitats, such as S. biflora which can be found some {{convert|4000|m}} above sea level in the Alps, or the East Greenland saxifrage (S. nathorstii). The genus is also abundant in the Eastern and Western Himalayan alpine shrub and meadows. Though the archetypal saxifrage is a small plant huddling between rocks high up on a mountain, many species do not occur in such a habitat and are larger (though still rather delicate) plants found on wet meadows. Various Saxifraga species are used as food plants by the caterpillars of some butterflies and moths, such as the Phoebus Apollo (Parnassius phoebus).[8] Charles Darwin – erroneously believing Saxifraga to be allied to the sundew family (Droseraceae) – suspected the sticky-leaved round-leaved saxifrage (S. rotundifolia), rue-leaved saxifrage (S. tridactylites) and Pyrenean saxifrage (S. umbrosa) to be protocarnivorous plants, and conducted some experiments whose results supported his observations,[9] but the matter has apparently not been studied since his time. CultivationNumerous species and cultivars of saxifrage are cultivated as ornamental garden plants, valued particularly as groundcover or as cushion plants in rock gardens and alpine gardens. Many require alkaline or neutral soil to thrive.[10] S. × urbium (London pride), a hybrid between Pyrenean saxifrage (S. umbrosa) and St. Patrick's cabbage (S. spathularis), is commonly grown as an ornamental plant.[2] Another horticultural hybrid is Robertsoniana saxifrage (S. × geum), derived from kidney saxifrage (S. hirsuta) and Pyrenean saxifrage.{{citation needed|date=October 2011}} Some wild species are also used in gardening. Cambridge University Botanic Garden hosts the United Kingdom's national collection of saxifrages.[2]Award of Garden MeritThe following species and cultivars have gained the Royal Horticultural Society's Award of Garden Merit:-[11] {{div col|colwidth=15em}}
UsesPurple saxifrage (S. oppositifolia) is a popular floral emblem. It is the territorial flower of Nunavut (Canada) and the county flower of County Londonderry in Northern Ireland.{{citation needed|date=October 2011}} Known as {{lang|no|rødsildre}} ("red saxifrage") in Norway, it also is the county flower of Nordland.{{citation needed|date=October 2011}} It is on the seal of Fitchburg State University, whose motto is "Perseverantia" in reference to the rock-breaking abilities of the plant over time. Tsukuba in Japan has as its city flower hoshizaki-yukinoshita (Katakana: ホシザキユキノシタ), the aptera form of Creeping saxifrage (S. stolonifera).{{citation needed|date=October 2011}} The leaves of the Japanese variety "yukinoshita" (literally "Under the snow") can also been eaten, and is consumed at least within the large southern island of Kyushu. It is prepared by frying the younger succulent leaves in tempura batter.{{citation needed|date=October 2011}} Selected species{{div col|colwidth=24em}}
Formerly placed herePlants formerly placed in Saxifraga are mainly but not exclusively Saxifragaceae. They include:{{citation needed|date=October 2011}}
Other "saxifragous" plantsSeveral plant genera have names referring saxifrages although they might not be close relatives of Saxifraga. They include:{{citation needed|date=October 2011}}
Some plants refer to Saxifraga in their generic names or specific epithets, either because they are also "rock-breaking" or because they resemble members of the saxifrage genus:{{citation needed|date=October 2011}}
References1. ^1 {{cite web |publisher=United States Department of Agriculture |date=February 9, 2005 |work=Germplasm Resources Information Network |url=http://www.ars-grin.gov/cgi-bin/npgs/html/genus.pl?10791 |title=Saxifraga L. |accessdate=January 20, 2009}} 2. ^1 2 3 {{cite web |url=http://www.botanic.cam.ac.uk/Botanic/Plant.aspx?p=27&ix=11&pid=2763&prcid=4&ppid=2763 |title=Saxifraga |work=National Plant Collections |publisher=Cambridge University Botanic Garden |accessdate=October 3, 2011}} 3. ^Roger Spencer, ed. Horticultural Flora of South-Eastern Australia. UNSW Press, 2002. p. 81. {{ISBN|9780868401676}} 4. ^{{cite book |author1=D. A. Webb |author2=R. J. Gornall |lastauthoramp=yes |year=1989 |title=Saxifrages of Europe |publisher=Christopher Helm |isbn=0-7470-3407-9 |page=19}} 5. ^{{cite journal |author=Douglas E. Soltis, Robert K. Kuzoff, Elena Conti, Richard Gornall & Keith Ferguson |year=1996 |title=matK and rbcL gene sequence data indicate that Saxifraga (Saxifragaceae) is polyphyletic |journal=American Journal of Botany |volume=83 |issue=3 |pages=371–382 |doi=10.2307/2446171}} 6. ^Flora of China 7. ^Flora of North America 8. ^{{cite book |author=Ivo Novák |year=1980 |title=A Field Guide in Colour to Butterflies and Moths |publisher=Octopus Books |isbn=0-7064-1293-1}} 9. ^{{cite book |author=Charles Darwin |year=1875 |chapter=Drosophyllum – Roridula – Byblis – glandular hairs of other plants – concluding remarks on the Droseraceae |title=Insectivorous Plants |edition=1st |pages=332–367 |publisher=J. Murray |location=London |url=https://archive.org/details/insectivorousplant00darw}} 10. ^1 {{cite book|title=RHS A-Z encyclopedia of garden plants|year=2008|publisher=Dorling Kindersley|location=United Kingdom|isbn=1405332964|pages=1136}} 11. ^{{cite web | url = https://www.rhs.org.uk/plants/pdfs/agm-lists/agm-ornamentals.pdf | title = AGM Plants - Ornamental | date = July 2017 | page = 95 | publisher = Royal Horticultural Society | accessdate = 1 November 2018}} 12. ^{{cite web | url = https://www.rhs.org.uk/Plants/328870/i-Saxifraga-i-Angelina-Johnson-(-i-fortunei-i-)-(5)/Details | title = RHS Plantfinder - Saxifraga ‘Angelina Johnson’ | accessdate = 1 November 2018}} 13. ^{{cite web | url = https://www.rhs.org.uk/Plants/135777/i-Saxifraga-i-Blackberry-and-Apple-Pie-(-i-fortunei-i-)-(5)/Details | title = RHS Plantfinder - Saxifraga ‘Blackberry and Apple Pie’ | accessdate = 1 November 2018}} 14. ^{{cite web|title=RHS Plant Selector - Saxifraga callosa|url=http://apps.rhs.org.uk/plantselector/plant?plantid=4110|accessdate=2 June 2013}} 15. ^{{cite web | url = https://www.rhs.org.uk/Plants/195374/i-Saxifraga-i-Conwy-Snow-(-i-fortunei-i-)-(5)/Details | title = RHS Plantfinder - Saxifraga ‘Conwy Snow’ | accessdate = 1 November 2018}} 16. ^{{cite web | url = https://www.rhs.org.uk/Plants/179269/i-Saxifraga-i-Coolock-Kate-(7)/Details| title = RHS Plantfinder - Saxifraga ‘Coolock Kate’ | accessdate = 1 November 2018}} 17. ^{{cite web | url = https://www.rhs.org.uk/Plants/104323/i-Saxifraga-i-Cumulus-(7)/Details| title = RHS Plantfinder - Saxifraga ‘Cumulus’ | accessdate = 1 November 2018}} 18. ^{{cite web|title=RHS Plant Selector - Saxifraga fortunei|url=http://apps.rhs.org.uk/plantselector/plant?plantid=1795|accessdate=2 June 2013}} 19. ^{{cite web | url = https://www.rhs.org.uk/Plants/71455/i-Saxifraga-i-Gregor-Mendel-(×-i-apiculata-i-)-(7)/Details| title = RHS Plantfinder - Saxifraga ‘Gregor Mendel’ (× fortunei) | accessdate = 1 November 2018}} 20. ^{{cite web | url = https://www.rhs.org.uk/Plants/51081/i-Saxifraga-i-Lagraveana-(-i-paniculata-i-)-(8)/Details | title = RHS Plantfinder - Saxifraga ‘Lagraveana’ | accessdate = 1 November 2018}} 21. ^{{cite web|title=RHS Plant Selector - Saxifraga 'Lutea'|url=http://apps.rhs.org.uk/plantselector/plant?plantid=4112|accessdate=2 June 2013}} 22. ^{{cite web|title=RHS Plant Selector - Saxifraga 'Minor'|url=http://apps.rhs.org.uk/plantselector/plant?plantid=4114|accessdate=2 June 2013}} 23. ^{{cite web | url = https://www.rhs.org.uk/Plants/328882/i-Saxifraga-i-Moe-(-i-fortunei-i-)-(5)/Details| title = RHS Plantfinder - Saxifraga ‘Moe’ | accessdate = 1 November 2018}} 24. ^{{cite web | url = https://www.rhs.org.uk/Plants/204222/i-Saxifraga-i-Monarch-(8)/Details| title = RHS Plantfinder - Saxifraga ‘Monarch’ | accessdate = 1 November 2018}} 25. ^{{cite web | url = https://www.rhs.org.uk/Plants/96144/i-Saxifraga-i-Mount-Nachi-(-i-fortunei-i-)-(5)/Details | title = RHS Plantfinder - Saxifraga ‘Mount Nachi’ | accessdate = 1 November 2018}} 26. ^{{cite web | url = https://www.rhs.org.uk/Plants/195379/i-Saxifraga-i-Peach-Melba-(7)/Details| title = RHS Plantfinder - Saxifraga ‘Peach Melba’ | accessdate = 1 November 2018}} 27. ^{{cite web | url = https://www.rhs.org.uk/Plants/172923/i-Saxifraga-i-(Silver-Farreri-Group)-Reginald-Farrer-(8)/Details | title = RHS Plantfinder - Saxifraga (Silver Farreri Group) ‘Reginald Farrer’ | accessdate = 1 November 2018}} 28. ^{{cite web | url = https://www.rhs.org.uk/Plants/89829/i-Saxifraga-i-Rokujo-(-i-fortunei-i-)-(5)/Details| title = RHS Plantfinder - Saxifraga ’Rokujo’ (fortunei) | accessdate = 1 November 2018}} 29. ^{{cite web|title=RHS Plant Selector - Saxifraga 'Rosea'|url=http://apps.rhs.org.uk/plantselector/plant?plantid=4115|accessdate=2 June 2013}} 30. ^{{cite web | url = https://www.rhs.org.uk/Plants/328887/i-Saxifraga-i-Shiranami-(-i-fortunei-i-)-(5)/Details| title = RHS Plantfinder - Saxifraga ‘Shiranami’ | accessdate = 1 November 2018}} 31. ^{{cite web | url = https://www.rhs.org.uk/Plants/215621/i-Saxifraga-i-(Southside-Seedling-Group)-Slack-s-Ruby-Southside-(8)/Details| title = RHS Plantfinder - Saxifraga (Southside Seedling Group) ‘Slack’s Ruby Southside’ | accessdate = 1 November 2018}} 32. ^{{cite web | url = https://www.rhs.org.uk/Plants/92308/i-Saxifraga-i-(Silver-Farreri-Group)-Snowflake-(8)/Details | title = RHS Plantfinder - Saxifraga (Silver Farreri Group) ‘Snowflake’ | accessdate = 1 November 2018}} 33. ^{{cite web | url = https://www.rhs.org.uk/Plants/215667/i-Saxifraga-i-(Southside-Seedling-Group)-Southside-Star-(8)/Details | title = RHS Plantfinder - (Southside seedling Group) ‘Southside Star’ | accessdate = 1 November 2018}} 34. ^{{cite web|title=RHS Plant Selector - Saxifraga stolonifera|url=http://apps.rhs.org.uk/plantselector/plant?plantid=1797|accessdate=2 June 2013}} 35. ^{{cite web | url = https://www.rhs.org.uk/Plants/252010/i-Saxifraga-i-Sue-Drew-(-i-fortunei-i-)-(5)/Details| title = RHS Plantfinder - Saxifraga ‘Sue Drew’ | accessdate = 1 November 2018}} 36. ^{{cite web | url = https://www.rhs.org.uk/Plants/148163/i-Saxifraga-i-Font-Face-times-New-Roman-Sugar-Plum-Fairy-FONT-Toujya-(-i-fortunei-i-)-(5)/Details | title = RHS Plantfinder - Saxifraga {{tdes|Sugar Plum Fairy}}=‘Toujya’ | accessdate = 1 November 2018}} 37. ^{{cite web|title=RHS Plant Selector - Saxifraga 'Theoden'|url=http://apps.rhs.org.uk/plantselector/plant?plantid=4108|accessdate=2 June 2013}} 38. ^{{cite web|title=RHS Plant Selector - Saxifraga 'Tumbling Waters'|url=http://apps.rhs.org.uk/plantselector/plant?plantid=4117|accessdate=2 June 2013}} 39. ^{{cite web|title=RHS Plant Selector - Saxifraga × urbium|url=http://apps.rhs.org.uk/plantselector/plant?plantid=1798|accessdate=2 June 2013}} 40. ^{{cite web | url = https://www.rhs.org.uk/Plants/91882/i-Saxifraga-i-Venetia-(-i-paniculata-i-)-(8)/Details| title = RHS Plantfinder - Saxifraga ‘Venetia’ (paniculata) | accessdate = 1 November 2018}} 41. ^{{cite web | url = https://www.rhs.org.uk/Plants/86835/i-Saxifraga-i-Whitehill-(8)/Details| title = RHS Plantfinder - Saxifraga ‘Whitehill’ | accessdate = 1 November 2018}} 42. ^1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 Umberto Quattrocchi. CRC World Dictionary of Plant Names: Common Names, Scientific Names, Eponyms. Synonyms, and Etymology. CRC Press, 1999. p.2395-2396. {{ISBN|9780849326738}} 43. ^{{cite journal|last=Knaben|first=G.|title=Saxifraga osloensis n. sp., a tetraploid species of the Tridactylites section|journal=Nytt Magasin for Botanikk|year=1934|pages=117–138}} 44. ^Readers Digest Nature Lover's Library Wild Flowers of Britain, page 192, published 1988 External links{{Commons category|Saxifraga}}
3 : Saxifraga|Saxifragaceae genera|Garden plants |
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