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词条 Stachys
释义

  1. Description

  2. Circumscription

  3. Fossil record

  4. Diversity

      Formerly placed here  

  5. See also

  6. References

  7. External links

{{For|the first bishop of Byzantium|Stachys the Apostle}}{{Italic title}}{{taxobox
|name = Stachys
|image = 20130703Ziest Saarbruecken1.jpg
|image_caption = Stachys sylvatica
|regnum = Plantae
|unranked_divisio = Angiosperms
|unranked_classis = Eudicots
|unranked_ordo = Asterids
|ordo = Lamiales
|familia = Lamiaceae
|subfamilia = Lamioideae
|genus = Stachys
|genus_authority = L.[1]
|type_species = Stachys sylvatica
|type_species_authority = L.
|subdivision_ranks = Species
|subdivision = About 300; see text
|synonyms_ref=[2]
|synonyms = *Betonica L.
  • Galeopsis Hill 1756 not L. 1753 nor Adans. 1763 nor Moench 1794
  • Galeopsis Moench 1794 not L. 1753 nor Adans. 1763 nor Moench 1794 nor Hill 1756
  • Zietenia Gled.
  • Trixago Haller
  • Bonamya Neck.
  • Eriostomum Hoffmanns. & Link
  • Tetrahitum Hoffmanns. & Link
  • Eriostemum Steud
  • Olisia Spach
  • Ortostachys Fourr.
  • Trixella Fourr.
  • Aspasia E.Mey. ex Pfeiff.
  • Stachyus St.-Lag.
  • Lamiostachys Krestovsk.
  • Menitskia (Krestovsk.) Krestovsk.

}}

Stachys is one of the largest genera in the flowering plant family Lamiaceae.[3] Estimates of the number of species in the genus vary from about 300,[3] to about 450.[4] The type species for the genus is Stachys sylvatica.[5] Stachys is in the subfamily Lamioideae.[3] Generic limits and relationships in this subfamily are poorly known.

The distribution of the genus covers Europe, Asia, Africa, Australasia and North America. Common names include hedgenettle,[6] heal-all, self-heal, woundwort, betony, and lamb's ears. Wood betony, S. officinalis, was the most important medicinal herb to the Anglo-Saxons of early medieval England.

The Chinese artichoke (S. affinis), is grown for its edible tuber.[4] Several species are cultivated as ornamentals. Woolly betony (S. byzantina) is a popular decorative garden plant.

Stachys was named by Linnaeus in Species Plantarum in 1753.[7] The name is derived from the Greek word σταχυς (stachys), meaning "an ear of grain",[8] and refers to the fact that the inflorescence is often a spike. The name woundwort derives from the past use of certain species in herbal medicine for the treatment of wounds.

Stachys species are used as food plants by the larvae of some Lepidoptera species, including the moths Coleophora auricella, C. lineolea, and C. wockeella, all recorded on S. officinalis. They are also widely used by the European wool carder bee (Anthidium manicatum), which scrape the hairs from the plant in order to use them for building their nests.[9]

Description

Stachys is a genus of shrubs and annual or perennial herbs. The stems vary from {{convert|50|-|300|cm|in|-1|abbr=on}} tall, with simple, opposite, triangular leaves, {{convert|1|-|14|cm|1|abbr=on}} long with serrate margins. In most species, the leaves are softly hairy. The flowers are {{convert|1|to|2|cm|in|1|abbr=on}} long, clustered in the axils of the leaves on the upper part of the stem. The corolla is 5-lobed with the top lobe forming a 'hood', varying from white to pink, purple, red or pale yellow.

Circumscription

The distinction between Stachys and other genera is unclear and has varied from one author to another. In 2002, a molecular phylogenetic study showed that Stachys officinalis is not closely related to the rest of the genus.[10] This study also found six other genera to be embedded within Stachys as it is currently circumscribed. The embedded genera are Prasium, Phlomidoschema, Sideritis, Haplostachys, Phyllostegia, and Stenogyne.

Fossil record

Stachys pliocenica fossil seeds are known from Upper Miocene strata of Bulgaria and Pliocene strata of south-eastern Belarus. The fossil seeds are similar to the seeds of Stachys cretica.[11]

Diversity

Species include:[12]

{{Div col |colwidth=27em}}
  • Stachys affinis Bunge – Chinese artichoke
  • Stachys ajugoides Benth. – bugle hedgenettle
  • Stachys alabamica B.R. Keener & L.J. Davenp. – Alabama hedgenettle
  • Stachys albens A.Gray – white hedgenettle, whitestem hedgenettle
  • Stachys albicaulis
  • Stachys alopecuros
  • Stachys alpina – alpine woundwort
  • Stachys annua (L.) L. – annual woundwort
  • Stachys arvensis L. – staggerweed, field woundwort
{{clear}}
  • Stachys bullata Benth.
  • Stachys byzantina K.Koch – woolly betony, lamb's ear
  • Stachys candida
  • Stachys chamissonis Benth. – great hedge nettle, coastal hedge nettle
  • Stachys chrysantha
  • Stachys ciliata
  • Stachys citrina
  • Stachys clingmanii – Clingman's hedge nettle
  • Stachys coccinea Jacq.
{{clear}}
  • Stachys cooleyae Heller – Cooley's hedge nettle
  • Stachys corsica
  • Stachys cretica L.
  • Stachys discolor
  • Stachys ehrenbergii
  • Stachys floridana Shuttlew. ex Benth. – Florida hedgenettle, Florida betony
  • Stachys germanica L. – downy woundwort
  • Stachys glutinosa
  • Stachys hispida – rough hedge-nettle
  • Stachys hyssopifolia
  • Stachys iva
{{clear}}
  • Stachys lanata
  • Stachys lavandulifolia
  • Stachys libanotica
  • Stachys macrantha (K.Koch) Stearn
  • Stachys macrostachya
  • Stachys manantlanensis B.L.Turner
  • Stachys mexicana Benth. – Mexican hedge nettle
  • Stachys milanii
  • Stachys monnieri (Gouan) P.W.Ball
{{clear}}
  • Stachys nelsonii B.R. Keener & L.J. Davenp. – Nelson's hedgenettle
  • Stachys officinalis (L.) Trevis. – betony
  • Stachys palustris L. – marsh woundwort, marsh hedge-nettle
  • Stachys pumila
  • Stachys pycnantha Benth.
  • Stachys recta L. – yellow woundwort
  • Stachys riddellii
  • Stachys rigida Benth. – rough hedgenettle
{{clear}}
  • Stachys scardia
  • Stachys stricta
  • Stachys sylvatica L. – hedge woundwort
  • Stachys tenuifolia Willd. – smooth hedgenettle
{{Div col end}}

Formerly placed here

  • Agastache foeniculum (Pursh) Kuntze (as S. foeniculum Pursh)
  • Leonurus japonicus Houtt. (as S. artemisia Lour.)[12]

See also

  • {{Commons category-inline}}
  • {{Wikispecies-inline}}
{{clear}}

References

1. ^{{cite web |url = http://www.ars-grin.gov/cgi-bin/npgs/html/genus.pl?11452 |title = Genus: Stachys L. |work = Germplasm Resources Information Network |publisher = United States Department of Agriculture |date = 2006-11-03 |accessdate = 2010-11-11 |deadurl = yes |archiveurl = https://web.archive.org/web/20100529194302/http://www.ars-grin.gov/cgi-bin/npgs/html/genus.pl?11452 |archivedate = 2010-05-29 |df = }}
2. ^Kew World Checklist of Selected Plant Families
3. ^Harley, R. M., et al. 2004. "Labiatae". pages 167–275. In: Kubitzki, K. (editor) and J. W. Kadereit (volume editor). The Families and Genera of Vascular Plants volume VII. Springer-Verlag: Berlin; Heidelberg, Germany. {{ISBN|978-3-540-40593-1}}
4. ^Mabberley, D. J. 2008. Mabberley's Plant-Book third edition (2008). Cambridge University Press: UK.
5. ^Stachys In: Index Nominum Genericorum. In: Regnum Vegetabile (see External links below).
6. ^{{PLANTS|id=STACH|taxon=Stachys|accessdate=30 November 2015}}
7. ^Carolus Linnaeus. 1753. Species Plantarum 2:580. Laurentii Salvii. (see External Links below).
8. ^Umberto Quattrocchi. 2000. CRC World Dictionary of Plant Names volume I, page 91. CRC Press: Boca Raton; New York; Washington, DC;, USA. London, UK. {{ISBN|978-0-8493-2673-8}} (set). (see External links below).
9. ^{{cite journal | last1 = Eltz | first1 = Thomas | last2 = Küttner | first2 = Jennifer | last3 = Lunau | first3 = Klaus | last4 = Tollrian | first4 = Ralph | title = Plant secretions prevent wasp parasitism in nests of wool-carder bees, with implications for the diversification of nesting materials in Megachilidae | journal = Frontiers in Ecology and Evolution | date = 6 January 2015 | volume = 2 | doi = 10.3389/fevo.2014.00086}}
10. ^Lindqvist, C. and V. A. Albert. 2002. Origin of the Hawaiian endemic mints within North American Stachys (Lamiaceae). American Journal of Botany 89(10), 1709–24.
11. ^The Pliocene flora of Kholmech, south-eastern Belarus and it's correlation with other Pliocene floras of Europe by Felix Yu. VELICHKEVICH and Ewa ZASTAWNIAK - Acta Palaeobot. 43(2): 137–259, 2003
12. ^{{cite web | url = http://www.ars-grin.gov/cgi-bin/npgs/html/splist.pl?11452 | title = GRIN Species Records of Stachys | work = Germplasm Resources Information Network | publisher = United States Department of Agriculture | accessdate = 2010-11-11}}

External links

  • Stachys At:Index Nominum Genericorum At: References At: NMNH Department of Botany
  • [https://www.biodiversitylibrary.org/item/13830#22 Stachys] In: [https://www.biodiversitylibrary.org/page/358106#1 Species Plantarum] At: [https://www.biodiversitylibrary.org/ Biodiversity Heritage Library]
  • CRC World Dictionary of Plant Names: R-Z At: Botany & Plant Science At: Life Science At: CRC Press
{{Taxonbar|from=Q156294}}

2 : Lamiales genera|Stachys

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