释义 |
- Description
- Classification Uncertain or dubious taxa Misassigned taxa
- References Notes Bibliography
- External links
{{distinguish|Cyathium|Cyathus}}{{automatic taxobox | image = Cyathea-med3.jpg | image_caption = Cyathea medullaris growing in New Zealand | taxon = Cyathea | authority = J. E. Smith, 1793 | subdivision_ranks = Subgenera and sections | subdivision =- Subgenus Cyathea
- Subgenus Sphaeropteris
- Subgenus Hymenophyllopsis
| synonyms =- Acanthea Lindig, 1861
- Alsophila R. Brown, 1810
- Amphicosmia Gardner, 1842
- Chlamydia Moore, 1857
- Chnoophora Kaulfuss, 1824
- Cormiophyllum Newman, 1856
- Cyatheopsis Karsten, 1858
- Dichorexia Presl, 1847
- Dicranophlebia Martius, 1834
- Disphenia Presl, 1836
- Eatoniopteris Bommer, 1873
- Eleutheria Kunze, 1844
- Fourniera Bommer, 1873
- Gymnopremnon Lindig, 1861
- Gymnosphaera Blume, 1828
- Halophlebia Martius, 1834
- Hymenophyllopsis Goebel, 1929
- Nephelea Tryon, 1970
- Notocarpia Presl, 1836
- Notophoria Presl, 1848
- Sarcopholis
- Schizocaena W. J. Hooker, 1838
- Schizocaena J. E. Smith, 1838
- Sclephropteris P. G. Windisch
- Sphaeropteris Bernhardi, 1801
- Thysanobotrya Alderwerelt, 1918
- Trachypremnon Lindig, 1861
- Trichipteris Presl, 1822
- Trichopteris Spreng., orth. var.
}}Cyathea is a genus of tree ferns, the type genus of the fern order Cyatheales. The genus name Cyathea is derived from the Greek kyatheion, meaning "little cup", and refers to the cup-shaped sori on the underside of the fronds. DescriptionThe species of Cyathea are mostly terrestrial ferns, usually with a single tall stem. Rarely, the trunk may be branched or creeping. Many species also develop a fibrous mass of roots at the base of the trunk. The genus has a pantropical distribution, with over 470 species. They grow in habitats ranging from tropical rain forests to temperate woodlands. ClassificationThe classification below follows the findings of Conant et al. in 1996[1]. These workers concluded on molecular cpDNA and morphological evidence that a system of three clades – Alsophila, Cyathea and Sphaeropteris is the most accurate reflection of evolutionary lineages within the Cyatheaceae, Alsophila being the most basal and Cyathea and Sphaeropteris derived sister groups. - Subgenus
- //Cyathea subg. Cyathea">Cyathea
, section Alsophila {{div col|colwidth=18em}}- Cyathea abbottii
- Cyathea acanthophora
- Cyathea acrostichoides
- Cyathea acuminata
- Cyathea affinis
- Cyathea albidosquamata
- Cyathea alderwereltii
- Cyathea alleniae
- Cyathea alpicola
- Cyathea amboinensis
- Cyathea amintae
- Cyathea andersonii
- Cyathea aneitensis
- Cyathea annae
- Cyathea apiculata
- Cyathea apoensis
- Cyathea approximata
- Cyathea archboldii
- Cyathea ascendens
- Cyathea atropurpurea
- Cyathea australis
- Cyathea baileyana
- Cyathea balanocarpa
- Cyathea batjanensis
- Cyathea biformis
- Cyathea borneensis
- Cyathea × boytelii
- Cyathea brevipinna
- Cyathea brooksii
- Cyathea bryophila
- Cyathea buennemeijerii
- Cyathea callosa
- Cyathea camerooniana
- Cyathea capensis
- Cyathea catillifera
- Cyathea caudata
- Cyathea chinensis
- Cyathea christii
- Cyathea cincinnata
- Cyathea cinerea
- Cyathea coactilis
- Cyathea colensoi
- Cyathea × confirmis
- Cyathea corcovadensis
- Cyathea costalisora
- Cyathea costulisora
- Cyathea crassa
- Cyathea croftii
- Cyathea cucullifera
- Cyathea cunninghamii
- Cyathea cuspidata
- Cyathea dealbata
- Cyathea deckenii
- Cyathea decora
- Cyathea decrescens
- Cyathea dicksonioides
- Cyathea dimorpha
- Cyathea doctersii
- Cyathea dregei
- Cyathea × dryopteroides
- Cyathea edanoi
- Cyathea elongata
- Cyathea erinacea
- Cyathea eriophora
- Cyathea esmeraldensis
- Cyathea everta
- Cyathea excavata
- Cyathea exilis
- Cyathea fadenii
- Cyathea × fagildei
- Cyathea fenicis
- Cyathea ferruginea
- Cyathea foersteri
- Cyathea fulgens
- Cyathea fuliginosa
- Cyathea geluensis
- Cyathea gigantea
- Cyathea glaberrima
- Cyathea glabra
- Cyathea glaziovii
- Cyathea gleichenioides
- Cyathea gregaria
- Cyathea grevilleana
- Cyathea halconensis
- Cyathea hancockii
- Cyathea havilandii
- Cyathea henryi
- Cyathea heterochlamydea
- Cyathea hooglandii
- Cyathea hookeri
- Cyathea hornei
- Cyathea horridula
- Cyathea hotteana
- Cyathea humilis
- Cyathea hunsteiniana
- Cyathea hymenodes
- Cyathea imbricata
- Cyathea imrayana
- Cyathea incana
- Cyathea incisoserrata
- Cyathea inquinans
- Cyathea insulana
- Cyathea javanica
- Cyathea junghuhniana
- Cyathea kanehirae
- Cyathea kermadecensis
- Cyathea khasyana
- Cyathea klossii
- Cyathea latebrosa
- Cyathea latipinnula
- Cyathea ledermannii
- Cyathea lepidoclada
- Cyathea loerzingii
- Cyathea loheri
- Cyathea longipes
- Cyathea lurida
- Cyathea macarthurii
- Cyathea macgillivrayi
- Cyathea macgregorii
- Cyathea macropoda
- Cyathea madagascarica
- Cyathea magnifolia
- Cyathea manniana
- Cyathea marattioides
- Cyathea × marcescens
- Cyathea masapilidensis
- Cyathea media
- Cyathea × medinae
- Cyathea mesosora
- Cyathea metteniana
- Cyathea mexicana
- Cyathea microchlamys
- Cyathea microphylloides
- Cyathea mildbraedii
- Cyathea milnei
- Cyathea minor
- Cyathea modesta
- Cyathea mossambicensis
- Cyathea muelleri
- Cyathea negrosiana
- Cyathea nicklesii
- Cyathea nigrolineata
- Cyathea nigropaleata
- Cyathea nilgirensis
- Cyathea nockii
- Cyathea obtusiloba
- Cyathea ogurae
- Cyathea oinops
- Cyathea oosora
- Cyathea orientalis
- Cyathea pachyrrhachis
- Cyathea pallidipaleata
- Cyathea parva
- Cyathea patellifera
- Cyathea percrassa
- Cyathea perpelvigera
- Cyathea perpunctulata
- Cyathea perrieriana
- Cyathea physolepidota
- Cyathea plagiostegia
- Cyathea podophylla
- Cyathea polycarpa
- Cyathea polystichoides
- Cyathea portoricensis
- Cyathea pruinosa
- Cyathea pseudomuelleri
- Cyathea pubescens
- Cyathea punctulata
- Cyathea pycnoneura
- Cyathea raciborskii
- Cyathea ramispina
- Cyathea rebeccae
- Cyathea recommutata
- Cyathea recurvata
- Cyathea rigens
- Cyathea rubella
- Cyathea rubiginosa
- Cyathea rufopannosa
- Cyathea rupestris
- Cyathea saccata
- Cyathea salletii
- Cyathea salvinii
- Cyathea scandens
- Cyathea schlechteri
- Cyathea schliebenii
- Cyathea sechellarum
- Cyathea semiamplectens
- Cyathea serratifolia
- Cyathea setosa
- Cyathea setulosa
- Cyathea sinuata
- Cyathea smithii
- Cyathea solomonensis
- Cyathea spinulosa
- Cyathea srilankensis[2]
- Cyathea stuebelii
- Cyathea subdubia
- Cyathea subtripinnata
- Cyathea sumatrana
- Cyathea tenuicaulis
- Cyathea ternatea
- Cyathea thomsonii
- Cyathea tryoniana
- Cyathea tsilotsilensis
- Cyathea tussacii
- Cyathea urbanii
- Cyathea vandeusenii
- Cyathea vieillardii
- Cyathea walkerae
- Cyathea welwitschii
- Cyathea wengiensis
- Cyathea woodwardioides
- Cyathea woollsiana
- Cyathea zakamenensis
{{div col end}}- Subgenus
- //Cyathea subg. Cyathea">Cyathea
, section Cyathea {{div col|colwidth=18em}}- Cyathea acutidens
- Cyathea aemula
- Cyathea alata
- Cyathea albomarginata
- Cyathea alphonsiana
- Cyathea alstonii
- Cyathea amazonica
- Cyathea andina
- Cyathea arborea
- Cyathea armata
- Cyathea ars
- Cyathea aspera
- Cyathea atahuallpa
- Cyathea aterrima
- Cyathea atrovirens
- Cyathea barringtonii
- Cyathea × bernardii
- Cyathea bettinae
- Cyathea bicrenata
- Cyathea bipinnata
- Cyathea boliviana
- Cyathea borinquena
- Cyathea bradei
- Cyathea brevistipes
- Cyathea brunnescens
- Cyathea × calolepis
- Cyathea caracasana
- Cyathea cicatricosa
- Cyathea concordia
- Cyathea conformis
- Cyathea conjugata
- Cyathea corallifera
- Cyathea costaricensis
- †Cyathea cranhamii
- Cyathea cyatheoides
- Cyathea cyclodium
- Cyathea cystolepis
- Cyathea darienensis
- Cyathea decomposita
- Cyathea decorata
- Cyathea decurrens
- Cyathea delgadii
- Cyathea demissa
- Cyathea dichromatolepis
- Cyathea dissimilis
- Cyathea dissoluta
- Cyathea divergens
- Cyathea dombeyi
- Cyathea dudleyi
- Cyathea ebenina
- Cyathea estelae
- Cyathea falcata
- Cyathea frigida
- Cyathea fulva
- Cyathea furfuracea
- Cyathea gardneri{{efn|Cyathea gardneri (W. J. Hooker) Lellinger 1987, not C. gardneri W. J. Hooker 1844.}}
- Cyathea gibbosa
- Cyathea glauca
- Cyathea gracilis
- Cyathea halonata
- Cyathea harrisii
- Cyathea haughtii
- Cyathea hemiepiphytica
- Cyathea hirsuta
- Cyathea hodgeana
- Cyathea holdridgeana
- Cyathea howeana
- Cyathea impar
- Cyathea intramarginalis
- Cyathea jamaicensis
- Cyathea kalbreyeri
- Cyathea lasiosora
- Cyathea latevagens
- Cyathea lechleri
- Cyathea leucofolis
- Cyathea × lewisii
- Cyathea lockwoodiana
- Cyathea macrocarpa
- Cyathea macrosora
- Cyathea marginalis
- Cyathea microdonta
- Cyathea microphylla
- Cyathea microphylla{{efn|Cyathea microphylla (Klotzsch) Tryon 1970, not C. microphylla Mettenius 1856.}}
- Cyathea mucilagina
- Cyathea multiflora
- Cyathea multisegmenta
- Cyathea myosuroides
- Cyathea nanna
- Cyathea nesiotica
- Cyathea nigripes
- Cyathea nodulifera
- Cyathea notabilis
- Cyathea onusta
- Cyathea palaciosii
- Cyathea paladensis
- Cyathea pallescens
- Cyathea parianensis
- Cyathea parva{{efn|Cyathea parva (Maxon) Tryon 1976, not C. parva Copeland 1942.}}
- Cyathea parvula
- Cyathea pauciflora
- Cyathea petiolata
- Cyathea phalaenolepis
- Cyathea phalerata
- Cyathea phegopteroides
- Cyathea pilosissima
- Cyathea pinnula
- Cyathea platylepis
- Cyathea poeppigii
- Cyathea praecincta
- Cyathea pseudonanna
- Cyathea pubens
- Cyathea punctata
- Cyathea pungens
- Cyathea robertsiana
- Cyathea rufa
- Cyathea ruiziana
- Cyathea sagittifolia
- Cyathea schiedeana
- Cyathea schlimii
- Cyathea senilis
- Cyathea simplex
- Cyathea sipapoensis
- Cyathea speciosa
- Cyathea squamulosa
- Cyathea steyermarkii
- Cyathea stipularis
- Cyathea stokesii
- Cyathea stolzei
- Cyathea straminea
- Cyathea subtropica
- Cyathea suprastrigosa
- Cyathea surinamensis
- Cyathea tenera
- Cyathea tortuosa
- Cyathea trichiata
- Cyathea tryonorum
- Cyathea ursina
- Cyathea valdecrenata
- Cyathea venezuelensis
- Cyathea villosa
- Cyathea weatherbyana
- Cyathea wendlandii
- Cyathea werffii
- Cyathea williamsii
{{Div col end}}- Subgenus
- //Cyathea sect. Sphaeropteris">Sphaeropteris
, section Sphaeropteris {{div col|colwidth=18em}}- Cyathea aciculosa
- Cyathea aeneifolia
- Cyathea agatheti
- Cyathea albifrons
- Cyathea albosetacea
- Cyathea alternans
- Cyathea angiensis
- Cyathea angustipinna
- Cyathea aramaganensis
- Cyathea arthropoda
- Cyathea assimilis
- Cyathea atrospinosa
- Cyathea atrox
- Cyathea auriculifera
- Cyathea binuangensis
- Cyathea brackenridgei
- Cyathea brownii
- Cyathea brunei
- Cyathea brunoniana
- Cyathea capitata
- Cyathea carrii
- Cyathea celebica
- Cyathea contaminans
- Cyathea cooperi
- Cyathea crinita
- Cyathea cuatrecasassi
- Cyathea curranii
- Cyathea deminuens
- Cyathea discophora
- Cyathea elliptica
- Cyathea elmeri
- Cyathea feani
- Cyathea felina
- Cyathea fugax
- Cyathea fusca
- Cyathea gardneri
- Cyathea hainanensis
- Cyathea inaequalis
- Cyathea insignis
- Cyathea insularum
- Cyathea integra
- Cyathea intermedia
- Cyathea × lathamii
- Cyathea leichhardtiana
- Cyathea lepifera
- Cyathea leucolepis
- Cyathea leucotricha
- Cyathea lunulata
- Cyathea macrophylla
- Cyathea magna
- Cyathea marginata
- Cyathea medullaris
- Cyathea megalosora
- Cyathea mertensiana
- Cyathea microlepidota
- Cyathea moluccana
- Cyathea moseleyi
- Cyathea nigricans
- Cyathea novae-caledoniae
- Cyathea obliqua
- Cyathea obscura
- Cyathea papuana
- Cyathea parksii
- Cyathea parvipinna
- Cyathea persquamulifera
- Cyathea philippinensis
- Cyathea pilulifera
- Cyathea polypoda
- Cyathea princeps
- Cyathea procera
- Cyathea propinqua
- Cyathea pulcherrima
- Cyathea quindiuensis
- Cyathea robinsonii
- Cyathea robusta
- Cyathea rosenstockii
- Cyathea runensis
- Cyathea sarasinorum
- Cyathea senex
- Cyathea setifera
- Cyathea sibuyanensis
- Cyathea squamulata
- Cyathea stipipinnula
- Cyathea strigosa
- Cyathea subsessilis
- Cyathea suluensis
- Cyathea tenggerensis
- Cyathea teysmannii
- Cyathea tomentosa
- Cyathea tomentosissima
- Cyathea trichodesma
- Cyathea trichophora
- Cyathea tripinnata
- Cyathea tripinnatifida
- Cyathea truncata
- Cyathea vaupelii
- Cyathea verrucosa
- Cyathea vittata
- Cyathea wallacei
- Cyathea werneri
- Cyathea whitmeei
- Cyathea womersleyi
- Cyathea zamboangana
{{Div col end}}- Subgenus
- //Cyathea subg. Hymenophyllopsis">Hymenophyllopsis
{{div col|colwidth=32em}}- Cyathea asplenioides syn.: Hymenophyllopsis asplenioides
- Cyathea ctenitoides syn.: Hymenophyllopsis ctenitoides
- Cyathea dejecta syn.: Hymenophyllopsis dejecta
- Cyathea hymenophylloidessyn.: Hymenophyllopsis hymenophylloides
- Cyathea incognita syn.: Hymenophyllopsis incognita
- Cyathea tepuiana syn.: Hymenophyllopsis steyermarkii
- Cyathea trichomanoides syn.: Hymenophyllopsis superba
- Cyathea universitatis syn.: Hymenophyllopsis universitatis
{{div col end}}Uncertain or dubious taxa- Alsophila crassa Karsten, 1869
- Cyathea affinis M. Martens & Galeotti, 1842
Misassigned taxa{{div col|colwidth=32em}}- Cyathea abitaguensis (Domin) Domin, 1930 = Cnemidaria uleana
- Cyathea antillana Domin, 1930 = Cnemidaria grandifolia
- Cyathea aristata Domin, 1930 = Cnemidaria apiculata
- Cyathea choricarpa (Maxon) Domin, 1929 = Cnemidaria choricarpa
- Cyathea commutata Sprengel, 1804 = Cnemidaria horrida
- Cyathea decurrentiloba Domin, 1930 = Cnemidaria decurrens
- Cyathea elegantissima (Fée) Domin, 1929 = Cnemidaria decurrens
- Cyathea ewanii Alston, 1958 = Cnemidaria ewanii
- Cyathea grandifolia Willdenow, 1810 = Cnemidaria grandifolia
- Cyathea grandis (Maxon) Domin, 1929 = Cnemidaria mutica
- Cyathea guatemalensis (Maxon) Domin, 1929 = Cnemidaria decurrens
- Cyathea horrida (Linnaeus) J. E. Smith, 1793 = Cnemidaria horrida
- Cyathea imrayana (W. J. Hooker) Domin, 1929 = Cnemidaria grandifolia
- Cyathea karsteniana Klotzsch, 1852 = Cnemidaria karsteniana
- Cyathea kohautiana (Presl) Domin, 1929 = Cnemidaria grandifolia
- Cyathea leprieurii (Kunze) Domin, 1929 = Cnemidaria cruciata
- Cyathea liebmannii Domin, 1929 = Cnemidaria decurrens
- Cyathea lucida (Fée) Domin, 1929 = Cnemidaria decurrens
- Cyathea minita Kaulfuss, 1824 = Cnemidaria grandifolia
- Cyathea mutica (Christ) Domin, 1929 = Cnemidaria mutica
- Cyathea obtusa (Kaulfuss) Domin, 1929 = Cnemidaria grandifolia
- Cyathea roraimensis (Domin) Domin, 1929 = Cnemidaria roraimensis
- Cyathea spectabilis (Kunze) Domin, 1929 = Cnemidaria spectabilis
- Cyathea spectabilis var. colombiensis Stolze, 1974 = Cnemidaria spectabilis var. colombiensis
- Cyathea spectabilis var. longipinna (Domin) Domin, 1929 = Cnemidaria spectabilis var. spectabilis
- Cyathea spectabilis var. trinitensis (Domin) Domin, 1929 = Cnemidaria spectabilis var. spectabilis
- Cyathea subarborescens Domin, 1930 = Cnemidaria uleana
- Cyathea subincisa (Kunze) Domin, 1929 = Cnemidaria speciosa
{{div col end}}References1. ^{{cite book|last1=Conant|first1=D.S.|last2=Raubeson|first2=L.A.|last3=Attwood|first3=D.K.|last4=Perera|first4=S.|last5=Zimmer|first5=E.A.|last6=Sweere|first6=J.A.|last7=Stein|first7=D.B.|chapter=Phylogenetic and evolutionary implications of combined analysis of DNA and morphology in the Cyatheaceae|date=1996|editor-last1=Camus|editor-first1=J.M.|editor-last2=Gibby|editor-first2=M.|editor-last3=Johns|editor-first3=R.J.|title=Pteridology in perspective|publisher=Royal Botanic Gardens|location=Kew, U.K.|pages=231-248|isbn=9781900347099}} 2. ^Ranil, R.H.G., D.K.N.G. Pushpakumara, T. Janssen, D.S.A. Wijesundara & D.U.M.B. Dhanasekara 2010. Cyathea srilankensis Ranil: a new tree fern species from Sri Lanka. American Fern Journal 100(1): 39–44. {{DOI|10.1640/0002-8444-100.1.39}}
Notes{{notelist}}Bibliography- Christenhusz, M.J.M. (2009). New combinations and an overview of Cyathea subg. Hymenophyllopsis (Cyatheaceae). Phytotaxa 1: 37-42.
- Large, M.F. and J.E. Braggins (2004). Tree Ferns. Timber Press, Inc. {{ISBN|0-88192-630-2}}
- Lehnert, M. (2006). New species and records of tree ferns (Cyatheaceae, Pteridophyta) in the northern Andes. Organisms, Diversity & Evolution 6: 321-322, electronic supplement 13: 1-11.
- Lehnert, M. (2009). Three new species of scaly tree ferns (Cyathea-Cyatheaceae) from the northern Andes. Phytotaxa 2: 43-56.
- The International Plant Names Index
External links- Cyathea: World Species List
- Alsophila: World Species List
- Sphaeropteris: World Species List
- Smith's original description of the genus online at Project Gutenberg
- Community: Care and propagation of Treeferns (German/English)
- Fern Files: Cyathea
{{Commons category|position=left|Cyathea| Cyathea}}{{Taxonbar|from=Q121608}} 4 : Cyathea|Cyatheaceae|Fern genera|Pantropical flora |