| 词条 | Emuellidae | ||||||||
| 释义 |
| name = Emuellidae | fossil_range = {{fossilrange|517}} late Botomian | image = BalcoracaniaDailyi.png | image_caption = Balcoracania dailyi of the family Emuellidae Lower Cambrian Emu Shale Kangaroo Island, South Australia © Dave Simpson | regnum = Animalia | phylum = Arthropoda | classis = Trilobita | ordo = Redlichiida | subordo = Redlichiina | superfamilia = Emuelloidea | familia = Emuellidae | familia_authority = Pocock, 1970 | subdivision_ranks = Genera | subdivision =
}}Emuellidae are a small family of trilobites, a group of extinct marine arthropods, that lived during the late Lower Cambrian (late Botomian) of the East Gondwana supercontinent, in what are today South-Australia and Antarctica. Emuellidae can be recognized among trilobites in having a set of unique features. The headshield or cephalon has large genal spines reaching back as far as the 3rd to 6th segment of the thorax. The eye-ridges contact the back of the frontal lobe of the glabella and extend laterally and backwards, roughly parallel to the frontal and lateral rim of the cephalon. There are small, clearly incised pits at the junction between the eye-ridge and the frontal lobe of the cephalic axis (or glabella). The thorax reaches its greatest width at the 6th segment. The frontal part or prothorax consists of 6 segments, with number 5 and 6 fused, and the 6th carrying very large trailing spines. The rear part or opistothorax consists of a variable but extremely large number of seqments (up to 97).[1] Description(See the Trilobite article for a definition of morphological terms) Cephalon: Cranidium subquadrate, glabella cylindrical, slightly contracted at S3, three pairs of glabellar furrows, preglabellar field short or absent, eye ridge wide, long, directed slightly postero-laterally, palpebral lobe cresentic, posterior area of fixigena with fulcrum, free cheeks (or librigenae) with long spines; hypostome conterminant, attached to a narrow rostral plate. The thorax is divided into a prothorax of six segments (the 6th carrying very large, trailing, pleural spines and extremely long opisthothorax of up to 97 segments (Balcoraciana dailyi holds the record for greatest number of thoracic segments in a trilobite species). Pygidium: A minute, segmented disc. Taxonomic historyFossils now assigned to the Emuellidae were first discovered by Dr. B. Daily, of the Geology Department, University of Adelaide in 1956.[2] Position of the Emuellidae within the RedlichiidaOriginally, the Emuellidae were described as part of the Redlichiina. The primitive features prompted the theory that the Emuellids actually represented the stem group of all trilobites, with the Olenellina having secondary fused facial sutures.[2] Later, the Emuellidae were placed in their own superfamily (Emuelloidea), recognizing that trilobites lacking facial sutures (i.e. the Olenellina) are the stem group.[3] This was followed by the clustering of the Emuellidae in a new superfamily Ellipsocephaloidea.[4] Most recently, the Emuellidae are regarded an early branch of the Redlichiina suborder, the closest relatives being the genera Bigotina, Abadiella, and the close-knit group of the families Estaingiidae, Ichangiidae and Ellipsocephalidae.[1] Genera assigned to the EmuellidaeHolyoakia has previously been assigned to the Emuellidae.[5] The tailshiel (or pygidium) in Holyoakia is about the same size as its cranidium, with a well-defined axis, eight axial rings, well-developed pleural ribs and furrows, and a spiny margin. The pygidia of Emuella and Balcoracania however are poorly differentiated, minute, and have a smooth margin. Later scholars therefore placed Holyoakia in the Dorypygidae.[1]DistributionBalcoracania dailyi occurs in the late Lower Cambrian (late Botomian) of South Australia (White Point conglomerate, Cape d' Estaing and Emu Bay sections, Kangaroo Island;[6] Warragee Member, Billy Creek Formation, Flinders Range;[6][7] Coads Hill Member, Billy Creek Formation, Reaphook Hill).[8] Balcoracania sp. has been collected from the Lower Cambrian of Antarctica (Shackleton Limestone, central Transantarctic Mountains).[5]Emuella dalgarnoi is found in the late Lower Cambrian (late Botomian) of South Australia (Emu Bay Shale, Kangaroo Island).[6]Emuella polymera has been collected from the late Lower Cambrian (late Botomian) of South Australia (Cape d' Estaing section, Kangaroo Island).[6]Key to the species
References1. ^1 2 {{cite journal|last1=Paterson|first1=J.R.|last2=Edgecombe|first2=G.D.|year=2006|title=The Early Cambrian trilobite Family Emuellidae Popock, 1970: Systematic position and revision of Australian Species|journal=Journal of Paleontology|volume=85|pages=496–513|url= }} 2. ^1 {{cite journal|last=Pocock|first=K.J.|year=1970|title=The Emuellidae, a new family of trilobites from the Lower Cambrian of South Australia|journal=Palaeontology|volume=13|pages=522–562}} 3. ^{{cite journal|last1=Zhang|first1=W.T.|last2=Lu|first2=Y.H.|last3=Zhu|first3=Z.L.|last4=Qian|first4=Y.Y.|last5=Li|first5=H.L.|last6=Zhou|first6=Z.Y.|last7=Zhang|first7=S.G.|last8=Yuan|first8=J.L.|year=1980|title=Cambrian trilobite faunas of Southwestern China|journal=Palaeontologica Sinica|volume=159|pages=1–497}} 4. ^{{cite journal|last=Lin|first=T.R.|year=1990|title=Application of custer analysis to taxonomy of superfamilies and families of superorder Redlichiina (Trilobita)|journal=Acta Palaeontologica Sinica|volume=29|pages=94–105}} 5. ^1 {{cite journal|last1=Palmer|first1=A.R.|last2=Rowell|first2=A.J.|year=1995|title=Early Cambrian trilobites from the Shackleton Limestone of the Central Transantarctic Mountains|journal=Paleontological Society Memoir|volume=45|pages=1–28}} 6. ^1 2 3 {{cite journal|last=Popock|first=K.J.|year=1970|title=The Emuellidae, a new family of trilobites from the Lower Cambrian of South Australia|journal=Palaeontology|volume=13|pages=522–562}} 7. ^{{cite journal|last=Moore|first=P.S.|year=1979|title=Stratigraphy and depositional environments of the Billy Creek Formation (Cambrian), central and northern Flinders Range, South Australia|journal=Transactions of the Royal Society of South Australia|volume=103|pages=197–211}} 8. ^{{cite journal|last=Moore|first=P.S.|year=1980|title=Stratigraphy and depositional environments of the Billy Creek Formation (Cambrian), east of the Flinders Range, South Australia|journal=Transactions of the Royal Society of South Australia|volume=104|pages=117–132}} External links{{refbegin}}
| chapter = Burgess Shale-type Localities: The global picture | author = Hagadorn, J.W. | editor = Bottjer, D.J. | editor2 =W. Etter | editor3 =J.W. Hagadorn | editor4 =C.M. Tang | title=Exceptional Fossil Preservation -- A Unique View on the Evolution of Marine Life | publisher=Columbia University Press | year=2002 | isbn =0231102542
|author1=Greg Edgecombe |author2=the Australian Museum |lastauthoramp=yes | title=Australian Trilobites: A Species List and Bibliography | url=http://www.austmus.gov.au/palaeontology/research/trilobites03.htm? | accessdate=August 23, 2005
| author=Sam Gon III | title=A guide to the Orders of Trilobites | url=http://www.trilobites.info/index.htm | accessdate=August 23, 2005
| author=Nedin, C. | title=The Emu Bay Shale, a Lower Cambrian fossil Lagerstätte, Kangaroo Island, South Australia | journal=Memoirs of the Association of Australasian Palaeontologists | year=1995 | volume=18 | pages=31–40
|author = Simpson, Dave |title = Trilobites of South Australia |url = http://info.esc.net.au/~dasimpson/index.htm |accessdate = August 23, 2005 |deadurl = yes |archiveurl = https://web.archive.org/web/20060219192556/http://info.esc.net.au/~dasimpson/index.htm |archivedate = February 19, 2006 |df = }}{{refend}}
3 : Cambrian trilobites|Emuelloidea|Trilobite families |
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