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词条 Csehbánya Formation
释义

  1. Geology

  2. Invertebrate paleofauna

  3. Vertebrate paleofauna

      Fish   Amphibians   Squamates    Crocodyliformes    Turtles   Pterosaurs  Non-Avian Dinosaurs  Aves 

  4. See also

  5. References

{{Infobox rockunit
| name = Csehbánya Formation
| image =
| caption =
| type = Geological formation
| age = Late Santonian {{Fossil range|85|84}}
| period = Santonian
| prilithology = Siltstone Sandstone Conglomerate
| otherlithology = Clay
| namedfor = Csehbánya, Hungary
| namedby =
| region = Europe
| country = {{Flag|Hungary}}
| coordinates = {{Coord|47.24|N|17.64|E|display=inline}}
| unitof =
| subunits =
| underlies = Jákó Marl Formation. Localized Unconformity with Eocene Iharkút Conglomerate Formation present in some areas
| overlies = Unconformity with Upper Triassic Main Dolomite Formation and Upper Cretaceous Nagytárkány Bauxite Formation
| thickness = Varies, but up to {{convert|100-150|m|ft|abbr=on}}
| extent =
| area = Transdanubian Mountains
| map =
| map_caption =
}}

The Csehbánya Formation is a geological formation in the Transdanubian Mountains of Veszprém County, Hungary. The formation dates to the Late Santonian (around 85 million years ago) of the Late Cretaceous. It represents a floodplain environment as opposed to the swampy lacustrine environment of the simultaneous Ajka Coal Formation, though there is complete overlap in terms of fauna. It underlies the Jákó Marl formation, and laterally transitions to the Ajka Coal Formation.[1]

Geology

The lithology of the unit is a cyclic variation of conglomerate, sandstone, variegated siltstone, clay and marl layers with some sporadic thin coal seams.[2] It it uncomfortably overlies the Late Triassic Main Dolomite Formation, which has deposits of the Upper Cretaceous Nagytárkány Bauxite Formation within deep karstic sinkholes in the formation. There is a basal conglomerate of dolomite clasts at the base of the Csehbánya formation. The main exposed portion of the formation is called the Iharkút locality, which is a disused bauxite quarry. At this location due to Paleogene uplift, it is uncomfortably overlain by the Lutetian Ikharkút Conglomerate. In the upper levels of the formation, there are sandstone lens beds present, which represent channel bodies. These are indicative of an anastomosing fluvial system. The exposure at this locality is between 100–150 m. Most of the fossils are found within the SZ-6 site at the locality, which is interpreted as a lag channel deposit

Invertebrate paleofauna

Amber is known from both the Ajka Coal and Csehbánya formations, and is commonly referred to as ajkait. However most of the arthropod inclusions in these are undescribed, only being shown in photographs,[3] With only two species of Ceratopogonids (biting midges) in the extant genus Leptoconops and the extinct genus Adelohelea being described.[4] A list of known taxa is given below.

Invertebrates of the Csehbánya Formation
Genus Species Location Stratigraphic position Abundance Notes Images
AdeloheleaA. magyaricaBiting midge
LeptoconopsL. clavaBiting midge, genus currently extant
ColeopteraIndeterminateSee figure 3 on website
DipteraIndeterminateSee figure 4 on website
HymenopteraIndeterminateSee figure 6 on website
NematoceraIndeterminateSee figure 7 on website

Vertebrate paleofauna

Fishes, amphibians, turtles, squamates, crocodilians, dinosaurs,[5] birds and pterosaurs present in Veszprém, Hungary, near the village of Németbánya in the Iharkút locality, an open-pit bauxite mine.[6][7] Other finds include Abelisauridae indet.,[8] Rhabdodontidae indet.,[7] Tetanurae indet.[8] Coprolites[8] and eggshells[9] are also known from the locality.

{{clear}}

Fish

Fish of the Csehbánya Formation[10]
Genus Species Location Stratigraphic position Abundance Notes Images
AtractosteusIndeterminateIharkút locality, Hungary."Jaw fragments, teeth, scales, vertebrae"Gar, genus extant
CoelodusIndeterminatePossible dentary/premaxilla, vomers, prearticulars, teeth and possible scales[11]Pycnodontiform
VidalamiinaeIndeterminate"Two vertebrae"
AmiidaeIndeterminateA left anterior ceratohyal and a vertebra
ElopiformesIndeterminate13 vertebrae
EllimmichthyiformesIndeterminateAt least two distinct taxa represented by four and one vertebraeClupeimorph
cf. SalmoniformesIndeterminatetwo vertebrae
AcanthomorphaIndeterminateThree vertebrae and 18 fin spines
TeleosteiIndeterminateTeeth, vertebrae and cycloid scales
ActinopterygiiIndeterminate9 distinct teeth morphotypes and some teeth bearing elements

Amphibians

Hungarobatrachus[14]H. szukacsi

Iharkút locality, Hungary.

Amphibians of the Csehbánya Formation
Genus Species Location Stratigraphic position Abundance Notes Images
Albanerpeton[12]IndeterminateIharkút locality, Hungary.Skull and jaw elementsAlbanerpetonid
Bakonybatrachus[13]B. fedoriIharkút locality, Hungary.Pelvic elementsDiscoglossid frog
Pelvic elements

Advanced frog.

PalaeobatrachidaeIndeterminateIharkút locality, Hungary.Postcranial remains
PelobatidaeIndeterminateIharkút locality, Hungary.Postcranial remains

Squamates

Squamates of the Csehbánya Formation
GenusSpeciesLocationStratigraphic positionAbundanceNotesImages
BicuspidonB. aff. hatzegiensis[15]Fragmentary mandibles, maxillaePolyglyphanodontid
Chromatogenys[16]C. tiliquoidesPartial right mandibleScincomorph
Distortodon[17]D. rhomboideusDentaries, maxillaPolyglyphanodontid
Pannoniasaurus[18]P. inexpectatusPartial skeletonMosasaur, also known from Ajka Coal Formation[1]
Pelsochamops[19]P. infrequensFragmentary dentary, dentary fragmentsChamopsiid
Scincomorpha[20]IndeterminateFragmentary dentariesAt least 3 distinct taxa, 4 mentioned in paper, but unsure if this counts Chromatogenys

Crocodyliformes

Crocodyliformes of the Csehbánya Formation
Genus Species Location Stratigraphic position Abundance Notes Images
Iharkutosuchus[21]I. makadii"Complete and partial skulls, mandibles, various cranial elements, teeth"Hylaeochampsid
Doratodon[22]D. carcharidens"Teeth, Jaw fragment"
?TheriosuchusIndeterminate"Teeth, cranial elements"
?AllodaposuchusIndeterminate"Teeth, jaw fragments, cranial elements"

Turtles

Turtles of the Csehbánya Formation
GenusSpeciesLocationStratigraphic positionAbundanceNotesImages
FoxemysF. trabanti"Complete and partial skulls, mandibles, vertebrae, pectoral and pelvic girdle elements, limb bones, plastron and carapax fragments"
KallokibotionIndeterminateShell fragments
DortokidaeIndeterminateShell fragments

Pterosaurs

Bakonydraco[23]B. Galaczi

Geographically present in Veszprém, Hungary.

Pterosaurs of the Csehbánya Formation
Genus Species Location Stratigraphic position Abundance Notes Images
"Premaxilla, complete mandible, 56 mandibular symphyses"Tapejarid
AzdarchidaeIndeterminate"Cervical vertebrae, pelvic girdle elements, limb bones"
PterodactyloideaIndeterminate"3 mandibular symphyses"

Non-Avian Dinosaurs

Ajkaceratops[24]A. kozmai[24]Hungarosaurus[26]H.tormai

Geographically present in Veszprém, Hungary.

Pneumatoraptor[27]P.fodori

Geographically present in Veszprém, Hungary.

MochlodonM. vorosi[28]
Dinosaurs of the Csehbánya Formation
Genus Species Location Stratigraphic position Abundance Notes Images
Fused premaxillae and rostral bones, with fragments of the maxillae, four predentaries.[24] Referred teeth.[25]Bagaceratopsid Ceratopsian
Several partial skeletonsStruthiosaurine Nodosaur
Single left scapulocoracoidParavian
Partial skull and postcraniaRhabdodontid
StruthiosaurusIndeterminateHumerus[29] and partial hip, and pelvic armor elements.[30]StruthiosaurineNodosaur
ParavesIndeterminate"Teeth, caudals, limb bones"
AbelisauridaeIndeterminateFemur[31] and pedal ungual phalanx[27]
TetanuraeIndeterminateMore than 50 teeth[27]
Sauropoda[32]IndeterminateSingle tooth

Aves

Bauxitornis[33]B. mindszentyae
Aves of the Csehbánya Formation
Genus Species Location Stratigraphic position Abundance Notes Images
Single left metatarsus

An avisaurid enantiornithine.

EnantornithineIndeterminateLimb bones

See also

{{Portal|Earth sciences|Paleontology|Dinosaurs}}
  • List of dinosaur-bearing rock formations

References

1. ^{{cite journal|last1=Ősi|first1=Attila|last2=Bodor|first2=Emese Réka|last3=Makádi|first3=László|last4=Rabi|first4=Márton|title=Vertebrate remains from the Upper Cretaceous (Santonian) Ajka Coal Formation, western Hungary|journal=Cretaceous Research|volume=57|year=2016|pages=228–238|issn=0195-6671|doi=10.1016/j.cretres.2015.04.014}}
2. ^{{cite journal|last1=Botfalvai|first1=Gábor|last2=Haas|first2=János|last3=Bodor|first3=Emese Réka|last4=Mindszenty|first4=Andrea|last5=Ősi|first5=Attila|year=2016|title=Facies architecture and palaeoenvironmental implications of the upper Cretaceous (Santonian) Csehbánya formation at the Iharkút vertebrate locality (Bakony Mountains, Northwestern Hungary)|journal=Palaeogeography, Palaeoclimatology, Palaeoecology|volume=441|pages=659–678|doi=10.1016/j.palaeo.2015.10.018|issn=0031-0182}}
3. ^{{cite web |url=http://magyardinoszaurusz.hu/en/research-of-cretaceous-ambers-and-their-inclusions/ |title=Research of Cretaceous ambers and their inclusions: Cretaceous ambers from Hungary |publisher= Magyar Dinoszaurusz |access-date=21 April 2018}}
4. ^{{cite journal|last1=Borkent|first1=Art|title=Upper and Lower Cretaceous biting midges (Ceratopogonidae: Diptera) from Hungarian and Austrian amber and the Koonwarra Fossil Bed of Australia|journal=Stuttgarter Beiträge zur Naturkunde (B) Geologie und Paläontologie|volume=249|year=1997|pages=1–10|url=https://www.biodiversitylibrary.org/page/30064743#page/151/mode/1up}}
5. ^Weishampel, David B; et al. (2004). "Dinosaur distribution (Late Cretaceous, Europe)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 588-593. {{ISBN|0-520-24209-2}}.
6. ^Makádi, L., Botfalvai, G., Ősi, A. 2006: Egy késő-kréta kontinentális gerinces fauna a Bakonyból: halak, kétéltűek, teknősök, gyíkok. Földtani Közlöny 136/4, pp. 487-502.
7. ^Ősi, A. & Rabi, M., 2006. Egy késő-kréta kontinentális gerinces fauna a Bakonyból II.: krokodilok, dinoszauruszok, pteroszauruszok és madarak (The Late Cretaceous continental vertebrate fauna from the Bakony Mountains II: crocodiles, dinosaurs (Theropoda, Aves, Ornithischia), pterosaurs). Földtani Közlöny, 136, 4, 503–526.
8. ^{{Cite journal|last=Segesdi|first=Martin|last2=Botfalvai|first2=Gábor|last3=Bodor|first3=Emese Réka|last4=Ősi|first4=Attila|last5=Buczkó|first5=Krisztina|last6=Dallos|first6=Zsolt|last7=Tokai|first7=Richárd|last8=Földes|first8=Tamás|date=June 2017|title=First report on vertebrate coprolites from the Upper Cretaceous (Santonian) Csehbánya Formation of Iharkút, Hungary|journal=Cretaceous Research|volume=74|pages=87–99|doi=10.1016/j.cretres.2017.02.010|issn=0195-6671}}
9. ^{{Cite journal|last=Prondvai|first=Edina|last2=Botfalvai|first2=Gábor|last3=Stein|first3=Koen|last4=Szentesi|first4=Zoltán|last5=Ősi|first5=Attila|date=March 2017|title=Collection of the thinnest: A unique eggshell assemblage from the Late Cretaceous vertebrate locality of Iharkút (Hungary)|journal=Central European Geology|volume=60|issue=1|pages=73–133|doi=10.1556/24.60.2017.004|issn=1788-2281}}
10. ^{{Cite journal|last=Szabó|first=Márton|last2=Ősi|first2=Attila|date=September 2017|title=The continental fish fauna of the Late Cretaceous (Santonian) Iharkút locality (Bakony Mountains, Hungary)|journal=Central European Geology|volume=60|issue=2|pages=230–287|doi=10.1556/24.60.2017.009|issn=1788-2281}}
11. ^{{Cite journal|last=Szabó|first=Márton|last2=Gulyás|first2=Péter|last3=Ősi|first3=Attila|date=April 2016|title=Late Cretaceous (Santonian) pycnodontid (Actinopterygii, Pycnodontidae) remains from the freshwater deposits of the Csehbánya Formation, (Iharkút, Bakony Mountains, Hungary)|journal=Annales de Paléontologie|volume=102|issue=2|pages=123–134|doi=10.1016/j.annpal.2016.04.001|issn=0753-3969}}
12. ^{{Cite journal|last=Szentesi|first=Zoltán|last2=Gardner|first2=James D.|last3=Venczel|first3=Márton|date=March 2013|title=Albanerpetontid amphibians from the Late Cretaceous (Santonian) of Iharkút, Hungary, with remarks on regional differences in Late Cretaceous Laurasian amphibian assemblages|journal=Canadian Journal of Earth Sciences|volume=50|issue=3|pages=268–281|doi=10.1139/e2012-024|issn=0008-4077|bibcode=2013CaJES..50..268S}}
13. ^{{Cite journal|last=Szentesi|first=Zoltán|last2=Venczel|first2=Márton|date=April 2012|title=A new discoglossid frog from the Upper Cretaceous (Santonian) of Hungary|journal=Cretaceous Research|volume=34|pages=327–333|doi=10.1016/j.cretres.2011.11.012|issn=0195-6671}}
14. ^{{Cite journal|author=Zoltán Szentesi and Márton Venczel |year=2010 |title=An advanced anuran from the Late Cretaceous (Santonian) of Hungary |url=http://www.schweizerbart.de/resources/downloads/paper_previews/75036.pdf |journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen |volume=256 |issue=3 |pages=291–302 |doi=10.1127/0077-7749/2010/0054 }}
15. ^{{Cite journal|last=Makádi|first=László|date=December 2006|title=Bicuspidon aff. hatzegiensis (Squamata: Scincomorpha: Teiidae) from the Upper Cretaceous Csehbánya Formation (Hungary, Bakony Mts)|journal=Acta Geologica Hungarica|volume=49|issue=4|pages=373–385|doi=10.1556/ageol.49.2006.4.5|issn=0236-5278}}
16. ^{{Cite journal|last=Makádi|first=László|last2=Nydam|first2=Randall L.|date=2014-12-23|title=A new durophagous scincomorphan lizard genus from the Late Cretaceous Iharkút locality (Hungary, Bakony Mts)|journal=Paläontologische Zeitschrift|volume=89|issue=4|pages=925–941|doi=10.1007/s12542-014-0253-1|issn=0031-0220}}
17. ^{{Cite journal|last=Makádi|first=László|date=November 2013|title=A new polyglyphanodontine lizard (Squamata: Borioteiioidea) from the Late Cretaceous Iharkút locality (Santonian, Hungary)|journal=Cretaceous Research|volume=46|pages=166–176|doi=10.1016/j.cretres.2013.08.001|issn=0195-6671}}
18. ^{{Cite journal|last=Makádi|first=László|last2=Caldwell|first2=Michael W.|last3=Ősi|first3=Attila|date=2012-12-19|title=The First Freshwater Mosasauroid (Upper Cretaceous, Hungary) and a New Clade of Basal Mosasauroids|journal=PLOS ONE|volume=7|issue=12|pages=e51781|doi=10.1371/journal.pone.0051781|issn=1932-6203|pmc=3526648|pmid=23284766|bibcode=2012PLoSO...751781M}}
19. ^{{Cite journal|last=Makádi|first=László|date=July 2013|title=The first known chamopsiid lizard (Squamata) from the Upper Cretaceous of Europe (Csehbánya Formation; Hungary, Bakony Mts)|journal=Annales de Paléontologie|volume=99|issue=3|pages=261–274|doi=10.1016/j.annpal.2013.07.002|issn=0753-3969}}
20. ^{{Cite journal|last=Botfalvai|first=Gábor|last2=Ősi|first2=Attila|last3=Mindszenty|first3=Andrea|date=January 2015|title=Taphonomic and paleoecologic investigations of the Late Cretaceous (Santonian) Iharkút vertebrate assemblage (Bakony Mts, Northwestern Hungary)|journal=Palaeogeography, Palaeoclimatology, Palaeoecology|volume=417|pages=379–405|doi=10.1016/j.palaeo.2014.09.032|issn=0031-0182}}
21. ^{{Cite journal|last=Ősi|first=Attila|last2=Clark|first2=James M.|last3=Weishampel|first3=David B.|date=2007-02-01|title=First report on a new basal eusuchian crocodyliform with multicusped teeth from the Upper Cretaceous (Santonian) of Hungary|journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen|volume=243|issue=2|pages=169–177|doi=10.1127/0077-7749/2007/0243-0169|issn=0077-7749}}
22. ^{{Cite journal|last=Rabi|first=Márton|last2=Sebők|first2=Nóra|date=October 2015|title=A revised Eurogondwana model: Late Cretaceous notosuchian crocodyliforms and other vertebrate taxa suggest the retention of episodic faunal links between Europe and Gondwana during most of the Cretaceous|journal=Gondwana Research|volume=28|issue=3|pages=1197–1211|doi=10.1016/j.gr.2014.09.015|issn=1342-937X|bibcode=2015GondR..28.1197R}}
23. ^{{Cite journal|last=Ösi|first=Attila|last2=Weishampel|first2=David B.|last3=Jianu|first3=Coralia M.|title=First evidence of azhdarchid pterosaurs from the Late Cretaceous of Hungary|journal=Acta Palaeontologica Polonica|volume=50|issue=4|year=2005|pages=777–787|url=https://www.app.pan.pl/article/item/app50-777.html}}
24. ^{{cite journal | last1 = Ősi | first1 = A. | last2 = Butler | first2 = R.J. | last3 = Weishampel | first3 = David B. | year = 2010 | title = A Late Cretaceous ceratopsian dinosaur from Europe with Asian affinities | url = | journal = Nature | volume = 465 | issue = 7297| pages = 466–468 | doi = 10.1038/nature09019 | pmid=20505726|bibcode = 2010Natur.465..466O }}
25. ^{{Cite journal|last=Virág|first=Attila|last2=Ősi|first2=Attila|date=2017-04-27|title=Morphometry, Microstructure, and Wear Pattern of Neornithischian Dinosaur Teeth From the Upper Cretaceous Iharkút Locality (Hungary)|journal=The Anatomical Record|volume=300|issue=8|pages=1439–1463|doi=10.1002/ar.23592|pmid=28371453|issn=1932-8486}}
26. ^Ösi, Attila. 2005. Hungarosaurus tormai, a new ankylosaur (Dinosauria) from the Upper Cretaceous of Hungary. Journal of Vertebrate Paleontology 25(2):370-383, June 2005.
27. ^Ősi, A., Apesteguía, S.M and Kowalewski, M. (In press). "Non-avian theropod dinosaurs from the early Late Cretaceous of Central Europe." Cretaceous Research, {{doi|10.1016/j.cretres.2010.01.001}}
28. ^{{Cite journal | last1 = Ősi | first1 = A. | last2 = Prondvai | first2 = E. | last3 = Butler | first3 = R. | last4 = Weishampel | first4 = D. B. | editor1-last = Evans | editor1-first = Alistair Robert | title = Phylogeny, Histology and Inferred Body Size Evolution in a New Rhabdodontid Dinosaur from the Late Cretaceous of Hungary | doi = 10.1371/journal.pone.0044318 | journal = PLoS ONE | volume = 7 | issue = 9 | pages = e44318 | year = 2012 | pmid = 23028518| pmc =3448614 |bibcode = 2012PLoSO...744318O }}
29. ^{{Cite journal|last=Ősi|first=Attila|last2=Prondvai|first2=Edina|date=August 2013|title=Sympatry of two ankylosaurs (Hungarosaurus and cf. Struthiosaurus) in the Santonian of Hungary|journal=Cretaceous Research|volume=44|pages=58–63|doi=10.1016/j.cretres.2013.03.006|issn=0195-6671}}
30. ^{{Cite journal|last=Ősi|first=Attila|last2=Pereda-Suberbiola|first2=Xabier|date=July 2017|title=Notes on the pelvic armor of European ankylosaurs (Dinosauria: Ornithischia)|journal=Cretaceous Research|volume=75|pages=11–22|doi=10.1016/j.cretres.2017.03.007|issn=0195-6671}}
31. ^{{Cite journal|last=Ősi|first=Attila|last2=Buffetaut|first2=Eric|date=January 2011|title=Additional non-avian theropod and bird remains from the early Late Cretaceous (Santonian) of Hungary and a review of the European abelisauroid record|journal=Annales de Paléontologie|volume=97|issue=1–2|pages=35–49|doi=10.1016/j.annpal.2011.07.001|issn=0753-3969}}
32. ^{{Cite journal|last=Ősi|first=Attila|last2=Csiki-Sava|first2=Zoltán|last3=Prondvai|first3=Edina|date=2017-06-12|title=A Sauropod Tooth from the Santonian of Hungary and the European Late Cretaceous 'Sauropod Hiatus'|journal=Scientific Reports|volume=7|issue=1|pages=3261|doi=10.1038/s41598-017-03602-2|issn=2045-2322|pmc=5468229|pmid=28607394|bibcode=2017NatSR...7.3261O}}
33. ^{{cite journal |authors=Gareth J. Dyke, Attila Ősi |year=2010 |title=A review of Late Cretaceous fossil birds from Hungary |journal=Geological Journal |volume=45 |issue=4 |pages=434–444 |doi=10.1002/gj.1209}}
{{clr}}{{Cretaceous Footer}}{{DEFAULTSORT:Csehbanya Formation}}

5 : Santonian Stage|Geology of Hungary|Cretaceous Europe|Paleontology in Hungary|Upper Cretaceous Series of Europe

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